Skeletomusculature of Scelionidae (Hymenoptera: Platygastroidea): head and mesosoma
Bibliographic record
Abstract
The skeletomusculature of the head and mesosoma of the parasitoid wasp family Scelionidae is reviewed. Representatives of 27 scelionid genera are examined together with 13 non-scelionid taxa for comparison. Terms employed for other groups of Hymenoptera are reviewed, and a consensus terminology is proposed. External characters are redescribed and correlated with corresponding apodemes, muscles and putative exocrine gland openings; their phylogenetic importance is discussed. 229 skeletal structures were termed and defined, from which 84 are newly established or redefined. 67 muscles of the head and mesosoma are examined and homologized with those present in other Hymenoptera taxa. The presence of the cranio-antennal muscle, an extrinsic antennal muscle originating from the head capsule, is unique for Scelionidae. The dorsally bent epistomal sulcus and the corresponding internal epistomal ridge extend to the anterior margin of the oral foramen, the clypeo-pleurostomal line is absent and the tentorium is fused with the pleurostomal condyle. The frontal ledge is present in those scelionid genera having the anterior mandibular articulation located on the lateral margin of the oral foramen. The ledge corresponds to the site of origin of the mandibular abductor muscle, which is displaced from the genal area to the top of the frons. The protractor of the pharyngeal plate originates dorsally of the antennal foramen in Scelionidae. All scelionid genera have a postgenal bridge developed between the oral and occipital foramina. The propleural arm is reduced, muscles originating from the propleural arm in other Hymenoptera are situated on other propectal structures in Scelionidae. The profurcal bridge is absent. The first flexor of the fore wing originates from the posteroventral part of the pronotum in Scelionidae and Vanhorniidae, whereas the muscle originatesfrom the mesopleuron in all other Hymenoptera. The netrion apodeme anteriorly limits the site of origin of the first flexor of the fore wing. Three types of netrion are described on the basis of the relative position of the netrion apodeme and the posterior pronotal inflection. The occlusor muscle apodeme is absent in basal Scelionidae, the fan-shaped muscle originates from the pronotum. In Nixonia the muscle originates posterior to the netrion apodeme. The skaphion apodeme crosses the site of origin of the longitudinal flight muscle. The lateral and dorsal axillar surfaces and the axillar carina are defined and described for the first time in Platygastroidea. The retractor of the mesoscutum is reported in Scelionidae and the variability of the muscle and corresponding skeletal structures within the family is described. The term sternaulus is redefined on the basis of the site of origin of the mesopleuro-mesobasalare muscle. The term speculum is adopted from Ichneumonidae and Cynipoidea taxonomy on the basis of the site of origin of the mesopleuro-mesofurcal muscle. The remnants of the mesopleural ridge, sulcus and mesopleural arm and pit and the putative border between the mesepisternum and mesepimeron is discussed. The mesopleural depressor of the mesotrochanter sensu Gibson 1985 originates from the anterior extension of the mesofurca and therefore the muscle is redefined and referred to in the present study as the lateral mesofurco-mesotrochanteral muscle. In Nixonia, Sparasion, Idris and Gryon both the lateral and median mesofurco-mesotrochanteral muscles are present. The lateral mesofurco-mesotrochanteral muscle is present in Platygastridae. The second flexor of the hind wing at least partly originates from the posteriorly delimited area of the mesopectus in Scelionidae similarly to some other Proctotrupomorpha and Chalcidoidea. The serial homology of this area and the netrion is discussed. The possible serial homology of the medially elevated area of the metanotum and mesoscutellum and the usage of the term metascutellum in Apocrita is discussed with the descriptions of correlated internal structures. The anterior metanotal wing process is located on the independent humeral sclerite in Scelionidae, similar to other Apocrita except Cynipoidea. The metanotal depressor of the metatrochanter originates from the humeral sclerite in Scelionidae as well as in some other Proctotrupoidea. The metapleuron is extended secondarily dorsally of the metapleural ridge and corresponding metapleural sulcus in Scelionidae. In Telenominae, Gryonini and Baeini the metafurca is located posteriorly on the metadiscrimenal lamella.
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How this classification was reachedexpand
Full frame machine prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. The Gemma side is a direct model label for every work in the frame, read from the title-only record. The Codex side is a classifier learned from the 10,348 direct Codex labels and calibrated to design-weighted sample rates; fields without enough sample support carry no Codex call. Candidate is the union of the two sides; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels.
Distilled classifier scores by category (both heads)
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.000 | 0.000 |
| Meta-epidemiology (narrow) | 0.000 | 0.000 |
| Meta-epidemiology (broad) | 0.000 | 0.000 |
| Bibliometrics | 0.001 | 0.001 |
| Science and technology studies | 0.000 | 0.000 |
| Scholarly communication | 0.000 | 0.000 |
| Open science | 0.000 | 0.000 |
| Research integrity | 0.000 | 0.000 |
| Insufficient payload (model declined to judge) | 0.001 | 0.000 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from itClassification
machine, unvalidatedMachine predicted; a candidate call from one source (direct Gemma or distilled Codex), not a consensus.
How this classification was reached, model by model and score by score, is at the end of the page under "How this classification was reached".