Rhamphomyia (Ctenempis) albopilosa Coquillett

Rhamphomyia (Ctenempis) albopilosa Coquillett (Figs 1–4, 62A) Rhamphomyia albopilosa Coquillett, 1900: 418. Type-locality: Berg Bay, Alaska, USA. Other references: Malloch, 1919: 47 (expedition results); Melander, 1928: 185 (catalogue); Melander, 1965: 461 (catalogue); Danks, 1981: 465 (arctic insects); Yang et al., 2007: 191 (catalogue). Rhamphomyia calvimontis Cockerell, 1916: 123. syn. nov. Type-locality: Baldy Mtn., Colorado, USA. Other references: Melander, 1928: 187 (catalogue); Melander, 1965: 462 (catalogue); Yang et al., 2007: 192 (catalogue). Rhamphomyia (Dasyrhamphomyia) wuorentausi Frey, 1922: 67. syn. nov. Type-locality: Dudinka, Krasnoyarskiy Territory, Russia. Rhamphomyia (Ctenempis) wuorentausi Frey: Frey, 1935: 3 (key and new subgenus); Frey, 1955b: 513 (revision). Rhamphomyia (Rhamphomyia) wuorentausi Frey: Chvála & Wagner, 1989: 292 (catalogue); Yang et al., 2007: 191 (catalogue); Shamshev, 2016: 84 (checklist). Notes on synonymy. The male terminalia of R. albopilosa and R. calvimontis are identical. They differ slightly on the basis of whitish setae on thorax and abdomen in the former compared to yellowish setae in the latter. Also legs generally with more numerous and longer setae in R. calvimontis, especially on the mid femur. The differences in setation are not considered to justify separate species recognition. Type material examined. Rhamphomyia albopilosa: LECTOTYPE (here designated in order to fix identity of the species) ♂ (Fig. 1A), labelled (Fig. 1B): “ Berg Bay [Alaska]/ 6–10’99”; “ Harriman Expedition ’99/ T. Kincaid Collector ”; “ Type / No. 5227/ U.S. N.M. [red label]”; “ Rhamphomyia / albopilosa/ Coq.”; “USNMENT/ 01288291 [data matrix code]” (USNM). PARATYPE: not examined [not present in USNM] (1 ♂, USNM). Rhamphomyia calvimontis: HOLOTYPE ♂ (Fig. 1C), labelled (Fig. 1D): “above timber/ line. Baldy / Mt. Col. / July 24. ’15./ (Cockerell) [hand written]”; “ALMelander/ Collection / 1961”; “ Rhamphomyia / calvimontis/ Ckll. TYPE. [hand written]”; “ HOLOTYPE / Rhamphomyia / calvimontis Cock. [red label]”; “USNMENT/ 01312581 [data matrix code]”; “ LECTOTYPE / Rhamphomyia albopilosa / Coquillett/ des. Sinclair & Saigusa 2018 [red label]” (USNM). Rhamphomyia wuorentausi: LECTOTYPE (here designated in order to fix identity of the species) ♂ (Fig. 2A), labelled (Fig. 2B): “Dudinka [Krasnoyarskiy Territory, Russia]”; “Wuorentaus”; “1584”; “Mus. Zool. H:fors/ Spec. typ. No. 4673/ Rhamph. wuo/ rentausi Frey”; “ LECTOTYPE / Rhamphomyia wuorentausi / Frey/ des. Sinclair & Saigusa 2018 [red label]” (MZH). PARALECTOTYPES: Same data as lectotype (1 ♀, MZH); Kantaika [= Khan- tayka, Krasnoyarskiy Territory, Russia], J. Sahlberg (1 ♀, MZH). Additional material examined. CANADA. Northwest Territories: Aklavik, 28.vi.1956, E.F. Cashman (1 ♂, CNC); Aklavik, 29.vi.–3.vii.1956, R.E. Leech (1 ♂, 2 ♀, CNC); Masik Riv., Banks Is., 3–29.vii.1968, W. R.M. Mason (16 ♂, 9 ♀, CNC); Masik Riv., Banks Is., 7–30.vii.1968, G.E. Shewell (16 ♂, 21 ♀, CNC); Reindeer De- pot, Mackenzie Delta, 10.vii.1948, J. R. Vockeroth (1 ♀, CNC); - 21 mi E Tuktoyaktuk, 20.vi.–21.vii.1971, D.M. Wood (10 ♂, 9 ♀, CNC); Victoria Is., 71°17′N 114°W, 8–28.vii.1975, G. & M. Wood (2 ♀, CNC). Nunavut: Baker Lake, 2–23.vii.1947, T.N. Freeman (2 ♂, 4 ♀, CNC); Baker Lake, 6.viii.1949, R.G., (1 ♂, 1 ♀, CNC); Bathurst Inlet, 5–19.vii.1951, C.D. Bird (2 ♀, CNC); Bernard Harbour, 68°47′06″N 114°49′43″W, 10.vii.1987, bait trap (2 ♂, CNC); Bernard Harbour, Dolphin & Union Str., 3–10.vii.1988, J. Troubridge (1 ♂, UGIC); Fraser Lake, 68°45′N, 120°36′W, 19.vii.1969, G.E. Shewell (4 ♂, 4 ♀, CNC); Head of Clyde Inlet, Baffin Is., 7.viii.1958, G.E. Shewell (9 ♂, 6 ♀, CNC); same data, J.E.H. Martin, (1 ♂, 6 ♀, CNC); Cambridge Bay, 12.viii.1959, J.A. Downes (1 ♀, CNC); Cambridge Bay, 69.1207°, -105.42582°, MT, mesic, 11–15.vii.2011, NBP field party, (25 ♂, LEM); Cambridge Bay, 69.12157°, -105.43124°, MT, 15–19.vii.2011, mesic, Repl.3, NBP field party (69 ♂, LEM); same data except, CCDB-21421-D01, CCDB-21421-E06 (1 ♂, 1 ♀, barcoded, LEM); Cambridge Bay, 69.12177°, - 105.41688°, sweep, 13–15.vii.2011, wet, Repl.1, NBP field party (5 ♂, LEM); Cockburn Point, Canadian Arctic Ex- pedition, 3.ix.1914, F. Johansen (1 ♀, CNC); Coppermine, 30.vi.–20.vii.1951, S.D. Hicks (4 ♀, CNC); Coral Har- bour, Southampton Is., 3–17.vii.1948, G.E. Shewell (27 ♂, 48 ♀, CNC); Coral Harbour, 15–16.viii.1959, W. R.N. Mason (2 ♂, 2 ♀, CNC); Coral Harbour, 16.viii.1959, J.A. Downes (1 ♀, CNC); Eskimo Point, 12–13.vii.1950, G.G. DiLabio (4 ♀, CNC); Eskimo Point, 17.vii.1950, G. R. Roberts (1 ♂, CNC); Muskox Lk., 64°45′N, 108°10′W, 11–18.vii.1953, J.G. Chillcott, (1 ♂, 2 ♀ CNC); Padley [Padlei], 24–27.vii.1950, R.E. Duckworth (2 ♂, 3 ♀, CNC); Padley, 1.viii.1950, R.A. Hennigar (1 ♀, CNC); Ross Bay, Melville Peninsula, N 66°51′ W 84°53′, 19–22.vii.1948, G.E. Shewell; Cody & McArthur (3 ♀, CNC); Spence Bay, 14.vii.1951, J.G. Chillcott (1 ♀, CNC). Quebec: Sugluk [Salluit], 14–22.vii.1954, H. Huckel (2 ♂, 4 ♀, CNC). Yukon: British Mts. Firth River, 27.vii.1956, R.E. Leech (1 ♂, CNC); British Mts., 69°15′N 140°02′W, 25.vi.1984, G. & M. Wood & D. Lafontaine (1 ♀, CNC); Dempster Hwy, mi 87, 27.vi.–12.vii.1973, G. & D.M. Wood (4 ♀, CNC); Dempster Hwy, km 141, 22–24.vi.1982, G. & M. Wood (1 ♂, 1 ♀, CNC); Firth River, 13–17.vii.1956, E.F. Cashman (3 ♀, CNC); Firth River, 13–14.vii.1956, R.E. Leech (1 ♂, 3 ♀, CNC); Herschel Is., 11–30.vii.1953, C.D. Bird (11 ♂, 23 ♀, CNC); Herschel Is., 24–29.vii.1971, D.M. Wood (5 ♂, 2 ♀, CNC); Herschel Is., 29–30.vii.1971, W. R.M. Mason (2 ♀, CNC); Herschel Is., 29.vii.1916, F. Johansen, Canadian Arctic Expedition (1 ♀, CNC); Herschel Is., 5–26.vii.1953, J.S. Waterhouse (9 ♂, 11 ♀, CNC); North Fork Crossing, mi 42, Peel Plt. Rd. 3500 ft, 24.vi.1962, P.J. Skitsko (1 ♀, CNC); North Fork Cross- ing, mi 42, Peel Plt. Rd. 3500 ft, 6.vi.1962, R.E. Leech (1 ♀, CNC); Whitehorse, 6.vii.1988, F. Brodo (1 ♂, CNC). USA. Alaska: Arctic NWR: Atigun Gorge, 600–1100 m, 68.48931°N 149.20343°W, riverside MT, 16.vii.2002 (1 ♂, UAM); Cape Thompson, 23.vii.1961, R. Madge (1 ♂, CNC); Kotzebue, 26.vi.1951, R.I. Sailer (1 ♂, USNM); Nome Area, mi 55 Kougarok Rd., 65°05′23″N 164°40′20″W, 20–26.vi.2005, MT, J & R. Skevington (21 ♂, 6 ♀, CNC); Schrader Lake, 69°30′N, 145°W, 3–5.vii.1972, K.W. Philip (1 ♂, 1 ♀, CNC). Colorado: Chasm Lake, 12000 ft, 5.viii.1958, R.L. Parker (1 ♂, USNM); Fall River Pass, 12000 ft, 12.vii.1934 [collector?] (2 ♂, USNM); Pike’s Park, 14.vii.1901, Johnson (2 ♀, USNM); Summit L./ Flats, Mt. Evans, 12800 ft, 7–24.vii.1961, C.H. Mann (16 ♂, 25 ♀, CNC); Summit L., Mt. Evans, 12800 ft, 24.vii.1961, J.G. Chillcott (7 ♂, 2 ♀, CNC); Summit L., Mt. Evans, 12800 ft, Marshy lake and stream margin, 24.vii.1961, J.G. Chillcott (1 ♂, 2 ♀, CNC); Mt. Evans, Rocky Slope, 13600 ft, 4.viii.1961, J.G. Chillcott (2 ♂, 3 ♀, CNC); Summit L., Mt. Evans, 12800 ft, 24.vii.1961, S.M. Clark (1♂, 1♀, CNC); Summit L., Mt. Evans, 12800 ft, 24.vii.1961, B.H. Poole (6 ♂, 6 ♀, CNC); Mt. Evans, 13000 ft, on tundra, 28.vii.1961, C.H. Mann (2 ♂, 9 ♀, CNC); Mt. Evans, 12000 ft, 3.viii.1961, W. R.M. Mason (1 ♀, CNC); Mt. Evans, 13200 ft, 19.vii.1961, W. R.M. Mason (2 ♂, 4 ♀, CNC); Mt. Evans, 14000 ft, 25.vii.1961, J.G. Chillcott (2 ♀, CNC); Mt. Evans, 14000 ft, 25.vii.1961, W. R.M. Mason (1 ♂, 2 ♀, CNC); Mt. Evans, 14000 ft, 25.vii.1961, B.H. Poole (5 ♂, 7 ♀, CNC); Mt. Evans, 14000 ft, 27.vii.1961, J. R. Stainer (1 ♂, CNC); Mt. Evans, 13200 ft, 28.vii.1961, S.M. Clark (1 ♂, 2 ♀, CNC). Idaho: Juliaetta (2 ♂, 3 ♀, USNM). Washington: Pullman, 20.v.1917, A.L. Melander (4 ♂, 8 ♀, USNM). Diagnosis. This species is distinguished from other species of Ctenempis by the subshiny scutum, male sternite 6 with transverse cluster of stiff golden ventral setae and a pair of horn-like projections on male sternite 7. Females are without pennate setae on legs and wings are broader than males and more darkly infuscate. Redescription. Wing length 6.0– 6.8 mm. Male. Head dark in ground-colour, with greyish pruinescence on face, frons, postgena and occiput; oral margin black and shiny. Holoptic, eyes with ommatidia larger on upper half of eye, smaller on lower half. Frons represented by very small triangular space below ocellar tubercle and larger subtriangular space above antennae; short, curved setulae along eye margin from ocellar triangle to antenna. Margins of face slightly divergent. Ocellar triangle dark with numerous setae of variable lengths. Upper half of occiput bearing row of long, black postocular setae, subequal in length with ocellar setae; upper occipital setae black and long; lower occipital setae pale, silky, hair-like, as long as upper occipital setae and distinctly denser on postgena. Antenna dark; scape slightly more than 2X longer than pedicel; postpedicel nearly 3X longer than basal width; stylus nearly as long as scape. Palpus dark, bearing many long, dark setae. Clypeus reddish, base glossy with thin pruinescence apically; labrum dark and glossy, length 2X eye height; labellum dark and bearing many dark setae, shorter than palpal setae. Thorax dark with dense grey pruinescence on pleura, scutum somewhat subshiny; darker grey to black vittae between acr and dc rows extending to prescutellar depression. Pleura with orange tint along notopleural ridge, as well as on posterior margin of katepisternum; pleural sclerites with variable pronounced shadows of brown; anterior and posterior spiracles pale brown. Proepisternum and prosternum (Fig. 3D) with many long, pale to golden setae; similar setae on upper part of proepisternum in front of anterior spiracle. Antepronotum with cluster of long, pale, lateral hair-like setae; middle part with many short, pale setae. Postpronotal lobe clothed in long, hair-like, pale setae with 1 outstanding pprn. Mesonotal setae long, mostly hair-like, becoming stouter, darker and sparser posteriorly (southern specimens with all dark setae): acr biserial; dc and presut s

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