Nigrobaetis rhithralis

Nigrobaetis rhithralis (Soldán & Thomas, 1983) Figures 1–9, 10A–B, 11–60, Table 1 Baetis rhithralis Soldán & Thomas, 1983: Acta Entomol. Bohemoslov., 80: 356 ? Baetis rhithralis Soldán & Thomas, 1983: Novikova & Kluge, 1994: Entomol. Obozr., 53 (3): 632 [synonymy assumption; Nigrobaetis the subgenus of Baetis s.l.] Diphetor rhithralis (Soldán & Thomas), 1989 [sic!]: Waltz et al., 1994: Bull. Soc. Hist. Nat., Toulouse, 130: 34 [new generic combination] Diphetor rhithralis (Soldán & Thomas, 1983): El Alami et al., 2000: Zool. Baetisca, 11: 110 [faunistics, new record from Morocco] Nigrobaetis rhithralis (Soldán & Thomas, 1983): Thomas, 1998: Bull. Soc. Hist. Nat., Toulouse, 134: 17 [faunistics, formal new generic combination]; Zrelli et al., 2011: Rev. Suisse Zool., 118 (1): 7 [faunistics, new record from Tunisia]; Zrelli et al., 2012: 67; Zrelli et al., 2016: Inland Water Biology, Suppl. 3: 102 Nigrobaetis rhithralis (Soldán & Thomas): Lugo-Ortiz & de Moor, 2000: African Entomol., 8 (1): 72 [systematics; discussion on generic placement] Baetis rhithralis: Jacob, 2003: Lauterbornia, 47: 121 [species grouping; faunistics] Baetis rhithralis Soldán & Thomas, 1983: Schwarz-Waubke et al., 2003: Beitr. Naturk. Oberöster., 12: 409 [list of types] Baetis (Nigrobaetis) rhithralis Soldán & Thomas, 1983: Bauernfeind & Soldán 2012: The Mayflies of Europe (Ephemeroptera): 156, 574, fig. 69 [taxonomy, distribution, biology] Type series. Original description of the species N. rhithralis is based on the male imago (holotype), female subimago and larvae (paratypes). The type series consists of 16 larvae, 3 imagos of both sexes (2 male imagos including the holotype, and one female imago), as well as 7 subimagos of both sexes (3 males and 4 females). All adults were reared from larvae, with preserved subimaginal and larval skins. Genitalia, forelegs and hind wings of the holotype and one male imago paratype were mounted on slides in Canada balsam and Liquid de Faure; 3 larvae and 6 larval skins were mounted on slides in Euparal or Canada balsam. Other adults are partly damaged (see below). Type locality can be defined as follows: Algeria, Wilaya de Blida [Blida Province], Chréa National Park, Blidean Atlas (part of the Tell Atlas), Camp des Chênes, Gorge de la Chiffa, confluence of Oued Chiffa [Chiffa River] and Ruisseau des singes, 250–270 m a.s.l., 36°24'04"N, 2°45'50"E, 09.x.1981, T. Soldán leg. (see Figs 82, 83). In the original description (Soldán & Thomas 1983a), the authors erroneously indicated that the type material of N. rhithralis was collected in Oued Merdja. However, this stream (original name is Oued Merdja Kbira) is situated somewhat southern in upper part of Oued Chiffa. The same mistake was made with the type locality of B. (Rhodobaetis) chelif Soldán & Godunko, 2005, as a part of the paratypes (adults and larvae) of this species was collected in the same place and date as the type series of N. rhithralis (see Soldán et al. 2005). Investigated type material of N. rhithralis preserved in ethanol was originally labelled as follows: [Baetis rhithralis Sold. & Thomas T. Soldán det. 1982]; [ALGER, Oued Merdja, Rouisseau des Singes 9.10.81 T. Soldán]. Place of deposition: IE (T. Soldán collection). Other material: 1 larva, Tunisia, Bizerte, Joumine, Basin Ichkeul, Riv. Joumine (Upstream dam); 57°46’09’’N 9°31’29’’E; 06.iv.2009; coll. Sonia Zrelli (MZL collection). GBIFCH Code: GBIFCH00331449. Diagnosis. Nigrobaetis rhithralis can be distinguished from other representatives of the genus Nigrobaetis by the combination of following characters: male imago: facetted surface of turbinate eyes intensively dark orange, shaft with two ring of different colour [according to original description]; faceted surface weakly ellipsoidal; shaft moderately high and clearly diverge after middle of its length; hind wings with two longitudinal veins (Figs 7, 8); second longitudinal vein forked at 0.48–0.56 of its length; abdominal terga I, VII–X rusty dark brownish; unistyliger nearly square-shaped, without apico-internal process; segment II of forceps with small inner hump; segment III of forceps elongated and rounded apically; larva: surface of antennal segment I and II with scattered short setae (Fig. 49); flagellar segment with petalshaped scales and their sockets on surface, prominent slender spines on posterior margin; nearly U-shaped frontal suture (Fig. 50); labrum with 1+2 (sometimes up to 4) long submarginal setae (Figs 17, 18); subapical setae absent on glossae dorsally; segment III of labial palp oblong-shaped, with inner margin nearly straight or slightly concave; 9–12 stout setae with jagged margins along outer margin of femora, 22–26 on inner margin of tibia, and 14–25 on inner margin of tarsi; tarsal claw with 12–16 spines, without subapical tiny setae; posterior margin of abdominal terga ĪIII (rarely VI) with a row of relatively short, stout spines; surface of terga and legs (especially femora) with distinct dominance of petal-shaped scales; gill I absent, gills II–VI asymmetric, gill VII slightly asymmetric or nearly symmetric; posterior prolongation of paraproct absent; inner margin of paraproct and cercotractor with respectively 28–35 and 18–25 spines arranged in one row; transversal band absent on caudal filaments; terminal filament 1/2–2/3 of cerci; cuticular coloration of abdominal segments uniform, with a pair of paramedian pale strokes and spots. Description. Male imago: Measurements: body length 4.2 mm; forewing length 4.5̄ 4.6 mm; cercus length 5.0– 5.2 mm. Specimens are partly damaged. The proportions of turbinate eyes are given in Table 1; for details see also Novikova & Kluge (1994: 630, table 3). Due to long-term storage of the material in ethanol (more than 30 years), the colour pattern of male imagos and other adults of N. rhithralis slightly differs from those described by Soldán & Thomas (1983a) (Figs 1–4, 11, 12). Regarding the general colour pattern published in the original description, the body colour of historical material is faded. Cuticular coloration is poorly distinguishable, especially on abdominal segments. Head. Vast part unicolorous light brown; darker smudges seen proximally. Antennae yellowish basally, yellowish-white apically. Dirty yellowish smudge seen between lateral ocelli. Ocelli yellow to dirty yellow apically, black basally. Turbinate eyes moderately high, well separated and diverge near to middle of their length (Figs 2, 5, 6); distance between basal part of turbinate eyes approximated equal to width of central ocellus; distance between upper parts of eyes at least three times more than base of central ocellus (for other proportions see Table 1). Faceted surface of turbinate eyes ellipsoidal dorsally (Fig. 5), slightly flattened; outer and inner margins distinctly convex. Faceted surface unicolorous, fadedly dirty brown [intensively dark orange: original description]. Colour pattern of turbinate eyes shaft poorly preserved (Fig. 2), i.e. apical strip only slightly paler than basal strip [broad paler strip apically and dark brown strip basally: orig. descr.]. Thorax. General colour pitch brown; pro- and metathorax slightly paler than mesothorax, light yellowishbrown to brown (Fig. 1). Ventral side of thorax paler than dorsal side, yellowish-brown. Wings hyaline, translucent, unicolorous [milky grey: orig. descr.] (Fig. 1). Longitudinal venation of fore wings yellowish-white; cross venation pale, poorly visible; pterostigma slightly frosted-white, with 7–10 cross veins (several of them bifurcated). Hind wings elongated, 2.90–2.94 times longer than wide; costal process of hind wings well developed, not hooked (Figs 7, 8). Two well developed longitudinal veins present at hind wings, occasionally with short third vein near base of wing (Fig. 8). Second longitudinal vein forked at 0.48–0.56 of its length, with one intercalary vein in fork; second longitudinal vein fork asymmetrical. Legs yellowish to dirty yellowish, with slightly darker femora [whitish, unicolorous: orig. descr.]. Mean length ratio of tarsomeres of foreleg as in Table 1 and Fig. 9. Abdomen. Terga without visible pattern; segments II–VI uncoloured, translucent, with small greyish shadows anterolaterally [white, without markings: orig. descr.]; terga I, VII–X light brown, with inconspicuous light brown smudges [rusty dark brownish: orig. descr.]. Cerci unicoloured yellowish-grey, paler apically. Genitalia. General colour yellowish to light brown; styliger, including unistyliger segments darker, forceps segments paler. Proportions of individual forceps segments slightly differ from those in original description: unistyliger nearly square-shaped (Fig. 10A–B), only slightly tapered apically, without apico-internal process, approximately 0.78–0.82 times longer than wide; segment I conical, slightly narrowed towards apex [segment I oblong-shaped, about twice broader than long: orig. descr.]; segment II expanding apically, with small blunt hump on inner margin basally [segment II slightly tapered in middle, with inconspicuous bluntly pointed projection at the same place: orig. descr.]; segment III narrow and elongated, slightly expanding apically, rounded, 3.10–3.40 times longer than wide [2.5: orig. descr.]. Female imago. Measurements: body length 5.0 mm; forewing length 4.9̄5.0 mm; cercus broken. Specimen is partly damaged. General colour yellowish to yellowish brown, paler than in male imago (Fig. 3). Head. Yellow, with darker smudges near eyes dorsally. Posterior margin of head nearly straight, only slightly convex medially (Fig. 12). Antennae yellow to greyish-yellow, scape darker. Eyes widely separated, unicoloured black, ocelli whitish apically, black basally (Fig. 4). Thorax. Yellowish-brown to brown dorsally, yellow ventrally. Wings hyaline, translucent; forewings slightly frosted light yellow, with milky pterostigma (Fig. 3). Longitudinal veins of forewings yellow to whitish-yellow, paler in Cu–A field. Hind wings wit

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