The Photoreceptor Axon Projection and Its Evolution in the Neural Superposition Eyes of Some Primitive Brachyceran Diptera (Part 1 of 2)
Bibliographic record
Abstract
All known brachyceran flies have inherited the same trapezoidal pattern of rhabdomeres in the ommatidia of the compound eye, different from that of their presumed nematoceran ancestors. The projection pattern of axons to lamina cartridges needs to match this in order to generate efficient neural superposition vision. This pattern has been traced in serial sections or via stimulated dye uptake in two families that diverged from the dipteran stem line more than 200 million years ago (Rhagionidae, Stratiomyidae) and in one family of intermediate age (Dolichopodidae). All show the same asymmetric projection as modern muscoid families and a concordantly 180 degrees-twisted axon bundle, suggesting that evolution to the 'modern' versions of both optics and projection coincided closely in evolutionary time. The basic developmental plan of the cartridge has also been retained since the origin of the Brachycera, but the synaptic participation of at least one axon (that of photoreceptor R7) varies markedly between species.
Fetched live from OpenAlex and de-inverted. Abstracts are not stored in this database: the inverted indexes are 8.6 GB of the frame’s 9.3 GB of text, and the host has 13 GB free.
How this classification was reachedexpand
Full frame machine prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. The Gemma side is a direct model label for every work in the frame, read from the title-only record. The Codex side is a classifier learned from the 10,348 direct Codex labels and calibrated to design-weighted sample rates; fields without enough sample support carry no Codex call. Candidate is the union of the two sides; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels.
Distilled classifier scores by category (both heads)
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.000 | 0.000 |
| Meta-epidemiology (narrow) | 0.000 | 0.000 |
| Meta-epidemiology (broad) | 0.000 | 0.000 |
| Bibliometrics | 0.001 | 0.000 |
| Science and technology studies | 0.001 | 0.001 |
| Scholarly communication | 0.000 | 0.000 |
| Open science | 0.000 | 0.000 |
| Research integrity | 0.000 | 0.000 |
| Insufficient payload (model declined to judge) | 0.001 | 0.000 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from itClassification
machine, unvalidatedMachine predicted; a candidate call from one source (direct Gemma or distilled Codex), not a consensus.
How this classification was reached, model by model and score by score, is at the end of the page under "How this classification was reached".