Review of fungus gnats of the genus Tarnania Tuomikoski, with a phylogeny of the Rymosia s.l. genus group (Diptera: Mycetophilidae)
Bibliographic record
Abstract
Abstract The four known species of Tarnania Tuomikoski are reviewed, viz. T. dziedzickii (Edwards, 1941), T. fenestralis (Meigen, 1818), T. nemoralis (Edwards, 1941), and T. tarnanii (Dziedzicki, 1910). Males and females are re-described, photographed and their terminalia figured. An attempt is made to homologize and name specific substructures of the male gonostylus, widely recognizable within the tribe Exechiini. A phylogenetic analysis of the Tarnania species, including representatives of the six other genera of the Rymosia s. lat. genus group, strengthens the view that Tarnania is monophyletic. Tarnania is supported by six unambiguous character changes of which three are considered to be unique synapomorphies, viz. 1) lower parts of metepisternum and metepimeron covered with campaniform sensilla; 2) hind tibia with several curved bristles posterodorsally, placed irregularly in more than one row; and 3) anterior branch of male gonostylus forming an elongated, bifurcated lobe. The most parsimonious hypothesis for relationships among the species of Tarnania is ((T. tarnanii + T. nemoralis) + (T. dziedzickii + T. fenestralis)). The practice of regarding the genera of the Rymosia s. lat. genus group as an assemblage of plesiomorphic genera is also supported, the most parsimonious hypothesis for relationships among them being (Notolopha (Allodiopsis (Myrosia (Synplasta (Tarnania + (Rymosia + Pseudorymosia)))))). Tarnania is most diverse in northwestern Europe, with all four species known from Norway, Sweden, Great Britain, Belgium, Germany, Switzerland, and France. T. dziedzickii extends its distribution southwards to include the North African and the Near East Regions. T. fenestralis is widely distributed in the western Palaearctic Region and extends its distribution eastwards to the eastern Palaearctic region. Only T. tarnanii displays a wide Holarctic distribution including most of Russia, Alaska, Canada and Greenland.
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How this classification was reachedexpand
Full frame machine prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. The Gemma side is a direct model label for every work in the frame, read from the title-only record. The Codex side is a classifier learned from the 10,348 direct Codex labels and calibrated to design-weighted sample rates; fields without enough sample support carry no Codex call. Candidate is the union of the two sides; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels.
Distilled classifier scores by category (both heads)
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.000 | 0.001 |
| Meta-epidemiology (narrow) | 0.001 | 0.000 |
| Meta-epidemiology (broad) | 0.001 | 0.000 |
| Bibliometrics | 0.006 | 0.005 |
| Science and technology studies | 0.000 | 0.000 |
| Scholarly communication | 0.001 | 0.001 |
| Open science | 0.001 | 0.001 |
| Research integrity | 0.000 | 0.001 |
| Insufficient payload (model declined to judge) | 0.003 | 0.001 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from itClassification
machine, unvalidatedMachine predicted; a candidate call from one source (direct Gemma or distilled Codex), not a consensus.
How this classification was reached, model by model and score by score, is at the end of the page under "How this classification was reached".