Modes of extinction, pseudo-extinction and distribution in Middle Jurassic ammonites: terminology
Bibliographic record
Abstract
Mid-Jurassic Ammonitina (Cephalopoda, Mollusca) provide good examples of true and apparent "extinctions" (i.e., taxon or clade disappearances) at the local, regional, and global scales. A terminology is presented. Extinction is the termination of a phylogenetic lineage or entire clade (not of local demes or regional populations). Extinction was often preceded by progressive range contraction that resulted in diachronous regional disappearance ("extirpation") and occurred with the elimination of the last refuge. Other range contractions, however, were not terminal, but were followed by renewed expansions, resulting in temporary absence of the lineage over part of its known range only, due to pseudo-extinction. Some lineages, called Lazarus taxa, apparently disappeared entirely for short or extended periods by pseudotermination (causing a "phylogenetic hiatus"). This is an extreme form of pseudo-extinction with unknown refuge due to small size and (or) unsuitable facies and location. Lineage or clade reappearance may be in the form of new species, whose relationship to ancestral taxa has been problematic. Some disappearances can be explained with displacive competition, where the replacement taxon is either of endemic origin or an immigrant. Recent research in previously underexplored field areas has closed some of the gaps of documentation by finding the refuges. Range contractions and expansions, together with their regional disappearances and pseudo-extinctions, including pseudotermination, were often causally related to sea-level changes, especially eustasy. Most true extinctions, however, cannot be identified precisely, because they occurred in small populations and (or) refuges. Extinctions presumably did not coincide with global geoevents.
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How this classification was reachedexpand
Full frame machine prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. The Gemma side is a direct model label for every work in the frame, read from the title-only record. The Codex side is a classifier learned from the 10,348 direct Codex labels and calibrated to design-weighted sample rates; fields without enough sample support carry no Codex call. Candidate is the union of the two sides; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels.
Distilled classifier scores by category (both heads)
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.001 | 0.002 |
| Meta-epidemiology (narrow) | 0.000 | 0.000 |
| Meta-epidemiology (broad) | 0.000 | 0.000 |
| Bibliometrics | 0.003 | 0.002 |
| Science and technology studies | 0.001 | 0.002 |
| Scholarly communication | 0.001 | 0.001 |
| Open science | 0.001 | 0.001 |
| Research integrity | 0.000 | 0.000 |
| Insufficient payload (model declined to judge) | 0.001 | 0.000 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from itClassification
machine, unvalidatedMachine predicted; a candidate call from one source (direct Gemma or distilled Codex), not a consensus.
How this classification was reached, model by model and score by score, is at the end of the page under "How this classification was reached".