A phylogeny of the hemichordates based on morphological characters
Bibliographic record
Abstract
A comprehensive review of literature on all 15 genera constituting the phylum Hemichordata resulted in a morphological matrix of 105 characters. The echinoderms, tunicates, cephalochordates, and vertebrates were included in the analysis, and the cnidarians, polychaetes, and sipunculids were employed as outgroup taxa. The consensus tree supported the traditional view of a monophyletic Hemichordata, Echinodermata, Ambulacraria, and Chordata. The enteropneust families Spengelidae and Ptychoderidae were each monophyletic and sister-taxa, but there was no resolution among the family Harrimaniidae. A detailed sensitivity analysis provided (i) tree lengths of competing evolutionary hypothesis and (ii) a test of monophyly of groups under a variety of evolutionary models. It is argued that the ancestral deuterostome was a benthic vermiform organism with a terminal mouth and anus, well-developed circular and longitudinal muscles, a simple nerve plexus with little sign of regionalization, a pharynx with gill slits and collagenous gill bars, a cluster of vacuolated cells with myofilaments, produced iodotyrosine, and displayed direct development. The pterobranchs have lost many of these features as a consequence of evolving a small body size and living in tubes, but these features exist in present-day enteropneusts, suggesting that they are a plausible model for the proximate ancestor of deuterostomes.
Fetched live from OpenAlex and de-inverted. Abstracts are not stored in this database: the inverted indexes are 8.6 GB of the frame’s 9.3 GB of text, and the host has 13 GB free.
How this classification was reachedexpand
Full frame machine prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. The Gemma side is a direct model label for every work in the frame, read from the title-only record. The Codex side is a classifier learned from the 10,348 direct Codex labels and calibrated to design-weighted sample rates; fields without enough sample support carry no Codex call. Candidate is the union of the two sides; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels.
Distilled classifier scores by category (both heads)
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.000 | 0.001 |
| Meta-epidemiology (narrow) | 0.001 | 0.000 |
| Meta-epidemiology (broad) | 0.000 | 0.000 |
| Bibliometrics | 0.003 | 0.003 |
| Science and technology studies | 0.000 | 0.000 |
| Scholarly communication | 0.001 | 0.001 |
| Open science | 0.000 | 0.000 |
| Research integrity | 0.000 | 0.000 |
| Insufficient payload (model declined to judge) | 0.002 | 0.001 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from itClassification
machine, unvalidatedMachine predicted; a candidate call from one source (direct Gemma or distilled Codex), not a consensus.
How this classification was reached, model by model and score by score, is at the end of the page under "How this classification was reached".