Reconstructing a radiation: the chiton genus Mopalia in the north Pacific
Bibliographic record
Abstract
The chiton genus Mopalia Gray, 1847 is highly speciose despite showing little morphological differentiation. Many of the 24 extant species are conspicuous, large-bodied and ecologically important today, but pre-Pleistocene fossils for the genus are rare. Here, we use a combined analysis of four gene regions (16S and COI mtDNA, 18S and 28S rDNA) to estimate the phylogenetic relationships for Mopalia species and use the inferred phylogeny to analyse the group’s biogeography and patterns of speciation. We then use these molecular data to distinguish between two alternative interpretations of the fossil record, as there is a large temporal gap between the oldest fossils tentatively identified as Mopalia and the next oldest fossils (Miocene versus Plio-Pleistocene). Based on the estimated substitution rates from a wide variety of other marine animals, we conclude that the observed rates in Mopalia are consistent with a Miocene origin for the genus. Given this age for the group and assuming a molecular clock, most speciation events in Mopalia are inferred to have occurred on average ~5 Mya. The phylogenetic results indicate that most of the speciation events leading to extant species must have occurred along the western North American coast, though there appear to have been multiple spreading events across the Pacific. When considered along with results for the many other near-shore taxa that have similar distributions to Mopalia, our findings suggest the emergence of a coherent historical biogeography of the northern Pacific.
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How this classification was reachedexpand
Full frame machine prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. The Gemma side is a direct model label for every work in the frame, read from the title-only record. The Codex side is a classifier learned from the 10,348 direct Codex labels and calibrated to design-weighted sample rates; fields without enough sample support carry no Codex call. Candidate is the union of the two sides; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels.
Distilled classifier scores by category (both heads)
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.000 | 0.001 |
| Meta-epidemiology (narrow) | 0.000 | 0.000 |
| Meta-epidemiology (broad) | 0.000 | 0.000 |
| Bibliometrics | 0.002 | 0.001 |
| Science and technology studies | 0.001 | 0.001 |
| Scholarly communication | 0.001 | 0.001 |
| Open science | 0.000 | 0.001 |
| Research integrity | 0.001 | 0.001 |
| Insufficient payload (model declined to judge) | 0.001 | 0.000 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from itClassification
machine, unvalidatedMachine predicted; a candidate call from one source (direct Gemma or distilled Codex), not a consensus.
How this classification was reached, model by model and score by score, is at the end of the page under "How this classification was reached".