A revision of the Late Cretaceous–Paleocene eutherian mammal <i>Cimolestes</i> Marsh, 1889
Bibliographic record
Abstract
Cimolestes Marsh is a North American eutherian mammal primarily known from latest Cretaceous deposits in Alberta, Wyoming, Saskatchewan, and Montana. At present, five species of Cimolestes are considered valid, all Lancian in age; they include one of the largest North American Late Cretaceous therian mammals as well as one of the smallest, a size range far exceeding that within other genera of tribosphenic therians contemporary with Cimolestes, such as the leptictoid eutherian Gypsonictops Simpson or genera of alphadontid or pediomyid marsupials. Moreover, the species of Cimolestes display a disparity of dental morphology in addition to size well in excess of interspecific differences within these genera. Given these considerations, Cimolestes is clearly a grade-taxon, uniting species sharing an adaptively subzalambdodont dentition, but showing divergent specializations within this pattern that are inconsistent with monophyly of its presently included species. To correct these imbalances, this paper limits Cimolestes to Cimolestes incisus Marsh and Cimolestes stirtoni Clemens; Cimolestes magnus Clemens and Russell, Cimolestes cerberoides Lillegraven, and Cimolestes propalaeoryctes Lillegraven are reclassified in the new genera Altacreodus, Ambilestes, and Scollardius, respectively. Altacreodus magnus, having a massive shearing dentition, is reconfirmed as showing a relationship to some Tertiary ‘creodonts’ not shared by other species of Lancian cimolestids; Ambilestes cerberoides exhibits a distinctive molar wear pattern that emphasized horizontal grinding, not orthal shear; Scollardius propalaeoryctes, the smallest species in this revision and having hyper-faunivorous molars, was not ancestral to Paleogene Palaeoryctidae, as indicated in part by contradictions in premolar number and morphology.
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How this classification was reachedexpand
Full frame machine prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. The Gemma side is a direct model label for every work in the frame, read from the title-only record. The Codex side is a classifier learned from the 10,348 direct Codex labels and calibrated to design-weighted sample rates; fields without enough sample support carry no Codex call. Candidate is the union of the two sides; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels.
Distilled classifier scores by category (both heads)
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.001 | 0.002 |
| Meta-epidemiology (narrow) | 0.001 | 0.000 |
| Meta-epidemiology (broad) | 0.000 | 0.000 |
| Bibliometrics | 0.003 | 0.001 |
| Science and technology studies | 0.002 | 0.003 |
| Scholarly communication | 0.002 | 0.001 |
| Open science | 0.002 | 0.001 |
| Research integrity | 0.001 | 0.001 |
| Insufficient payload (model declined to judge) | 0.003 | 0.001 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from itClassification
machine, unvalidatedMachine predicted; a candidate call from one source (direct Gemma or distilled Codex), not a consensus.
How this classification was reached, model by model and score by score, is at the end of the page under "How this classification was reached".