Homolidae de Haan, 1839 and Homolodromiidae Alcock, 1900 (Crustacea: Decapoda: Brachyura) from the Pacific Northwest of North America and a reassessment of their fossil records
Bibliographic record
Abstract
New material collected from Cretaceous and Tertiary rocks of the Pacific Northwest of North America has prompted a reevaluation of the fossil record of the Homolidae de Haan, 1839 and the Homolodromiidae Alcock, 1900. The fossil records of the homolid generaHomolaLeach, 1815;HomolopsisBell, 1863; andHoplitocarcinusBeurlen, 1928 are restricted, andLatheticocarcinusBishop, 1988, which is synonymous withEohomolaCollins and Rasmussen, 1992 andMetahomolaCollins and Rasmussen, 1992, is reinstated as a distinctive genus. Thirteen new combinations resulted from reinstatement ofLatheticocarcinus: L. adelphinus(Collins and Rasmussen, 1992),L. affinis(Jakobsen and Collins, 1997),L. atlanticus(Roberts, 1962),L. brevis(Collins, Kanie, and Karasawa, 1992),L brightoni(Wright and Collins, 1972),L. centurialis(Bishop, 1992),L. declinata(Collins, Fraaye, and Jagt, 1995),L. dispar(Roberts, 1962),L. pikeae(Bishop and Brannen, 1992),L. punctatus(Rathbun, 1917),L. schlueteri(Beurlen, 1928),L. shapiroiBishop, 1988,L. spiniga(Jakobsen and Collins, 1997), andL. transiens(Segerberg, 1900). A new species,Latheticocarcinus ludvigseni, is described from Cretaceous rocks of British Columbia. The first fossil occurrence of the extant homolid genusParomolopsis, P. piersoninew species, is recorded from Miocene rocks of Oregon.Paromola pritchardiJenkins, 1977 is formally transferred toDagnaudusGuinot and Richer de Forges (1995) as suggested by Guinot and Richer de Forges (1995). The extinct family Prosopidae von Meyer, 1860 is referred to the Homolodromioidea Alcock, 1900, following previous work.Palehomola gorrelliRathbun, 1926 is transferred from the Homolidae to the Homolodromiidae, and the new genusRhinodromiais erected to containHomolopsis richardsoniWoodward, 1896, from Cretaceous rocks of British Columbia. A new terminology is suggested for describing the rostral area in homolodromiids, in an attempt to alleviate considerable confusion over that issue. The morphologic similarity of fossil and extant members in both the Homolidae and the Homolodromiidae suggest that these two brachyuran families are evolutionarily conservative, much as the lobsters are. In addition, the similar paleobiogeographic and evolutionary patterns seen in the two families suggests that either they are closely related or that brachyuran families exhibited similar evolutionary and dispersal trends early in their history.
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How this classification was reachedexpand
Full frame machine prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. The Gemma side is a direct model label for every work in the frame, read from the title-only record. The Codex side is a classifier learned from the 10,348 direct Codex labels and calibrated to design-weighted sample rates; fields without enough sample support carry no Codex call. Candidate is the union of the two sides; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels.
Distilled classifier scores by category (both heads)
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.000 | 0.000 |
| Meta-epidemiology (narrow) | 0.000 | 0.000 |
| Meta-epidemiology (broad) | 0.000 | 0.000 |
| Bibliometrics | 0.001 | 0.001 |
| Science and technology studies | 0.001 | 0.001 |
| Scholarly communication | 0.000 | 0.001 |
| Open science | 0.000 | 0.001 |
| Research integrity | 0.000 | 0.000 |
| Insufficient payload (model declined to judge) | 0.005 | 0.001 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from itClassification
machine, unvalidatedMachine predicted; a candidate call from one source (direct Gemma or distilled Codex), not a consensus.
How this classification was reached, model by model and score by score, is at the end of the page under "How this classification was reached".