Systematics, distributions and bionomics of the Catopocerini (eyeless soil fungivore beetles) of North America (Coleoptera: Leiodidae: Catopocerinae)
Bibliographic record
Abstract
This paper is a review and revision of the tribe Catopocerini (Coleoptera: Leoididae: Catopocerinae) of North America. It covers the following genera: Catopocerus Motschulsky, 1870 with five species east of the Mississippi River and the resurrected genus Pinodytes Horn, 1880 with 42 species in North America west of the Mississippi River. All species in the tribe are eyeless and wingless inhabitants of forest soil and litter. Larvae and adults probably feed on subterranean fungi. Pinodytes Horn is resurrected to valid generic status. A neotype is assigned for Catopocerus politus Motschulsky. Lectotypes are designated for Catops cryptophagoides (Mannerheim, 1852) (which is transferred to Pinodytes), and Pinodytes pusio Horn, 1892. The following new synonym is recognized: Catopocerus ulkei Brown, 1933 = Catopocerus politus Motschulsky, 1870. The 33 new species and their distributions are as follows: Pinodytes angulatus (NW Oregon, USA), P. borealis (central Alaska, USA), P. chandleri (N California, USA), P. colorado (Colorado, USA), P. constrictus (S California, USA), P. contortus (E California, USA), P. delnorte (NW California, USA), P. eldorado (E California, USA), P. fresno (central California, USA), P. garibaldi (NW Oregon, USA), P. gibbosus (S California, USA), P. haidagwaii (Haida Gwaii (formerly Queen Charlotte) Islands, British Columbia, Canada), P. humboldtensis (NW California, USA), P. idaho (NW Idaho, USA), P. isabella (N Idaho, USA), P. klamathensis (SW Oregon and NW California, USA), P. losangeles (S California, USA), P. marinensis (W California, USA), P. minutus (central California, USA), P. monterey ( SW California, USA), P. newtoni (Ozarks region to E Texas, USA), P. orca (SW Oregon, USA), P. parvus (NW California, USA), P. punctatus (W Idaho and E Washington, USA), P. sanjacinto (S California, USA), P. sequoia ( S central California, USA), P. setosus ( SW Oregon and NW California, USA), P. shasta (N California, USA), P. shoshone (N Idaho, USA), P. sinuatus (SW Oregon, USA), P. spinus (N central California, USA), P. tehama (N California, USA), and P. tuolumne (E central California, USA). The following new combinations are established: Pinodytes capizzii (Hatch, 1957), ex Catopocerus; P. cryptophagoides (Mannerheim, 1852), ex Catopocerus; P. imbricatus (Hatch, 1957), ex Catopocerus; P. newelli (Hatch, 1957), ex Catopocerus; P. ovatus (Hatch, 1957), ex Catopocerus; P. pusio Horn, 1892, ex Catopocerus; P. rothi (Hatch, 1957), ex Catopocerus; P. subterraneus (Hatch, 1935), ex Catopocerus; P. tibialis (Hatch, 1957), ex Catopocerus.
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How this classification was reachedexpand
Full frame machine prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. The Gemma side is a direct model label for every work in the frame, read from the title-only record. The Codex side is a classifier learned from the 10,348 direct Codex labels and calibrated to design-weighted sample rates; fields without enough sample support carry no Codex call. Candidate is the union of the two sides; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels.
Distilled classifier scores by category (both heads)
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.000 | 0.000 |
| Meta-epidemiology (narrow) | 0.000 | 0.000 |
| Meta-epidemiology (broad) | 0.000 | 0.000 |
| Bibliometrics | 0.001 | 0.002 |
| Science and technology studies | 0.000 | 0.000 |
| Scholarly communication | 0.001 | 0.001 |
| Open science | 0.000 | 0.000 |
| Research integrity | 0.000 | 0.000 |
| Insufficient payload (model declined to judge) | 0.001 | 0.000 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from itClassification
machine, unvalidatedMachine predicted; a candidate call from one source (direct Gemma or distilled Codex), not a consensus.
How this classification was reached, model by model and score by score, is at the end of the page under "How this classification was reached".