Evolution of Serial Patterns in the Vertebrate Pharyngeal Apparatus and Paired Appendages via Assimilation of Dissimilar Units
Bibliographic record
Abstract
Evolution of serially similar structures has attracted interest since the infancy of comparative morphology and embryology. A long-standing assumption is that the serial patterning reflects ancestral metamerism, which persists in preconceived character polarity from a primitive state of polyisomerism (a series of identical or similar units) to a derived state of anisomerism (a series of differentially specialized parts). We test this assumption against an alternative character polarity — from anisomeric to polyisomeric — in the vertebrate pharyngeal apparatus and paired appendages. We show that, contrary to what is usually assumed, serial similarity represents a derived state in both pharyngeal apparatus and paired appendages: the distinctly patterned structures secondarily assimilated each other. Acquisitions of serial similarity in the pharyngeal apparatus and paired appendages straddle major evolutionary events such as the origin of the jaw and fish-tetrapod transitions. We suggest that: (a) the origin of the jaw coincided with extension of the serial pharyngeal patterning onto the mandibular region; and (b) the pectoral and pelvic appendages have independent origins and their distal portions acquired serial similarities later during the fin-limb transitions.
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How this classification was reachedexpand
Full frame machine prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. The Gemma side is a direct model label for every work in the frame, read from the title-only record. The Codex side is a classifier learned from the 10,348 direct Codex labels and calibrated to design-weighted sample rates; fields without enough sample support carry no Codex call. Candidate is the union of the two sides; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels.
Distilled classifier scores by category (both heads)
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.000 | 0.001 |
| Meta-epidemiology (narrow) | 0.000 | 0.000 |
| Meta-epidemiology (broad) | 0.000 | 0.000 |
| Bibliometrics | 0.001 | 0.000 |
| Science and technology studies | 0.000 | 0.001 |
| Scholarly communication | 0.001 | 0.001 |
| Open science | 0.000 | 0.001 |
| Research integrity | 0.000 | 0.000 |
| Insufficient payload (model declined to judge) | 0.002 | 0.001 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from itClassification
machine, unvalidatedMachine predicted; a candidate call from one source (direct Gemma or distilled Codex), not a consensus.
How this classification was reached, model by model and score by score, is at the end of the page under "How this classification was reached".