Data from new taxa infer<i>Isoechiniscoides</i>gen. nov. and increase the phylogenetic and evolutionary understanding of echiniscoidid tardigrades (Echiniscoidea: Tardigrada)
Bibliographic record
Abstract
The marine heterotardigrades belonging to Echiniscoididae inhabit a wide range of substrates and represent the dominant tardigrade group in intertidal zones worldwide. Notably, the evolution of this family is interesting as it holds key evidence as to how heterotardigrades colonized terrestrial and freshwater environments. We inferred the phylogeny of Echiniscoididae using 28S and COI sequences from GenBank and from new specimens collected at Roscoff, France. Morphological characterizations of several major clades, inferred from the molecular data, are currently lacking. However, one of the clades is erected as a new taxon, Isoechiniscoidesgen. nov., for the former Echiniscoides higginsi-group, which comprises species with isonych claws, pillars in the epicuticle and exceptionally long sensory appendages. The type species for the new genus is described here as Isoechiniscoides sifaenov. gen., sp. nov.Isoechiniscoides comprises six clawed interstitial species, whereas Echiniscoides taxa possess a higher claw number and primarily inhabit algae, barnacles and lichens in tidal zones. The new genus, Isoechiniscoides, together with morphologically undetermined specimens from British Columbia (Canada), is sister-group to all other Echiniscoididae taxa. Consequently, we propose to establish Isoechiniscoidinae subfam. nov. to accommodate the new genus. Isoechiniscoidinae provides an evolutionary link between Echiniscoidea and Arthrotardigrada. The latter comprises exclusively marine and mostly interstitial species.
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How this classification was reachedexpand
Full frame machine prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. The Gemma side is a direct model label for every work in the frame, read from the title-only record. The Codex side is a classifier learned from the 10,348 direct Codex labels and calibrated to design-weighted sample rates; fields without enough sample support carry no Codex call. Candidate is the union of the two sides; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels.
Distilled classifier scores by category (both heads)
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.000 | 0.001 |
| Meta-epidemiology (narrow) | 0.001 | 0.000 |
| Meta-epidemiology (broad) | 0.000 | 0.000 |
| Bibliometrics | 0.002 | 0.002 |
| Science and technology studies | 0.001 | 0.001 |
| Scholarly communication | 0.001 | 0.001 |
| Open science | 0.000 | 0.000 |
| Research integrity | 0.000 | 0.001 |
| Insufficient payload (model declined to judge) | 0.003 | 0.002 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from itClassification
machine, unvalidatedMachine predicted; a candidate call from one source (direct Gemma or distilled Codex), not a consensus.
How this classification was reached, model by model and score by score, is at the end of the page under "How this classification was reached".