Osteology and Histology of Plesiosaurs (Reptilia: Sauropterygia) from Non-marine (Fluvial to Estuarine) and Marine Depositional Environments
Bibliographic record
Abstract
Plesiosaurs are a highly diverse group of secondarily-aquatic reptiles that lived from Late Triassic to Late Cretaceous time. Plesiosaurs are known primarily from marine deposits, but also occur sparingly in non-marine units. Here, I describe the osteology and histology of plesiosaurs from both non-marine (Dinosaur Park Formation, DPF) and marine (Bearpaw and Pierre Shale Formations; BF, PSF), Upper Cretaceous-aged deposits of western Canada. These sediments were deposited in the Western Interior Basin, which was inundated by a large marine corridor known as the Western Interior Seaway. Non-marine (fluvial to estuarine) sediments of the DPF have yielded a stratigraphically extensive collection of elasmosaurid plesiosaur specimens. These specimens are notable as they belong to individuals that are small-bodied relative to some elasmosaurid specimens from marine units. In this study, I test whether the small-bodiedness of DPF specimens is due to the presence of immature individuals or a small-bodied taxon. Analyses of select DPF elasmosaurid specimens indicate the presence of a small-bodied taxon. Furthermore, a well-preserved partial skeleton from this unit is recognized as the holotype of a new genus and species. The holotype and referred specimens span both estuarine and fluvial sediments and may indicate an example of niche-partitioning within the predominantly marine Plesiosauria. The small body size is also consistent with those of plesiosaurs from other non-marine units. I also describe select elasmosaurid and polycotylid specimens from the BF and PSF, respectively. One elasmosaurid belongs to a large-bodied taxon which may be closely-related to, or possibly conspecific with, the DPF taxon. Another elasmosaurid belongs to a small-bodied, likely new, taxon. The two polycotylids were originally considered to be a juvenile and adult of Dolichorhynchops bonneri, based on their similar morphologies and large discrepancy in size. However, both specimens are osteologically mature, possibly reflecting interspecific differences; alternatively, these body size differences may be due to sexual dimorphism. Finally, this dissertation constitutes one of the first histological studies on plesiosaurs from non-marine units and Canada. This study also contributes to our limited understanding of plesiosaur ontogeny.
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How this classification was reachedexpand
Full frame machine prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. The Gemma side is a direct model label for every work in the frame, read from the title-only record. The Codex side is a classifier learned from the 10,348 direct Codex labels and calibrated to design-weighted sample rates; fields without enough sample support carry no Codex call. Candidate is the union of the two sides; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels.
Distilled classifier scores by category (both heads)
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.000 | 0.000 |
| Meta-epidemiology (narrow) | 0.000 | 0.000 |
| Meta-epidemiology (broad) | 0.000 | 0.000 |
| Bibliometrics | 0.002 | 0.001 |
| Science and technology studies | 0.000 | 0.000 |
| Scholarly communication | 0.000 | 0.000 |
| Open science | 0.000 | 0.000 |
| Research integrity | 0.000 | 0.000 |
| Insufficient payload (model declined to judge) | 0.002 | 0.000 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from itClassification
machine, unvalidatedMachine predicted; a candidate call from one source (direct Gemma or distilled Codex), not a consensus.
How this classification was reached, model by model and score by score, is at the end of the page under "How this classification was reached".