Late Miocene eastwards transatlantic dispersal of flightless anchonine weevils (Coleoptera: Curculionidae: Molytinae)
Bibliographic record
Abstract
The weevil genera Aethiopacorep Voisin and Titilayo Cristóvão Lyal are the only native African members of the nearly pantropical and poorly known tribe Anchonini. All Anchonini are flightless, a trait likely limiting dispersal, yet these weevils are found on both sides of the Atlantic Ocean. A phylogenetic analysis of 79 terminals and 3248 aligned positions from one mitochondrial and two nuclear ribosomal fragments supports a clade of West African Anchonini nested within American Anchonini. As suggested by previous authors, the Asian genera Himalanchonus Zherikhin and Otibazo Morimoto do not form a clade with the tribe's core, and along with Cycloterinus Kolbe, Euthycodes Pascoe, Leptanchonus Morimoto, Nepalanchonus Zherikhin, and Tanyomus Champion, are here removed from Anchonini and placed as Molytinae incertae sedis. So defined, the monophyletic tribe Anchonini contains 36 genus-group names, all but two denoting American taxa. Using molecular clock analysis, we estimate the separation of the West African Anchonini from its American sister at 9.5-5.2 million years ago (Ma). This date greatly postdates the Cretaceous opening of the Atlantic Ocean (about 100 Ma) and, therefore, evokes a single transatlantic dispersal to West Africa, likely by over-water rafting, leading to subsequent diversification. We postulate this to be the first documented eastwards crossing of the Atlantic Ocean by terrestrial non-volant arthropods based on morphological and molecular data.
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How this classification was reachedexpand
Full frame distilled prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. Learned from the 10,348 direct Codex labels and 10,348 direct Gemma labels. Candidate is the union of thresholded teacher heads; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels or direct frontier model labels.
Codex and Gemma teacher scores by category
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.000 | 0.000 |
| Meta-epidemiology (narrow) | 0.000 | 0.000 |
| Meta-epidemiology (broad) | 0.000 | 0.000 |
| Bibliometrics | 0.000 | 0.000 |
| Science and technology studies | 0.000 | 0.000 |
| Scholarly communication | 0.000 | 0.000 |
| Open science | 0.000 | 0.000 |
| Research integrity | 0.000 | 0.000 |
| Insufficient payload (model declined to judge) | 0.001 | 0.000 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from itClassification
machine, unvalidatedMachine predicted; a candidate call from one teacher head, not a consensus.
How this classification was reached, model by model and score by score, is at the end of the page under "How this classification was reached".