Grylloptera - a unique origin of the stridulatory fi le in katydids, crickets, and their kin (Archaeorthoptera)
Why this work is in the frame
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Bibliographic record
Abstract
Topographic homology conjectures (= THCs) of male forewing venation in extant ensiferan orthopterans (crickets, katydids, and their kin) and their close stem-relatives are re-evaluated, in order to test competing hypotheses on the origin(s) of the fi le (a row of teeth located on the ventral side of the forewing and used in stridulation). A new set of THCs (= STHC) is proposed, based on morphological data on the species †zeuneri Sharov, 1968, obscura Walker, 1869, monstrosa Uhler, 1864, †madygenicus Sharov, 1968: p. 181, grandidieri de Saussure, 1877: p. 287, bimaculatus de Geer, 1773, villosiceps Chopard, 1951, frontalis Walker, 1869, gryllotalpa Linnaeus, 1758, vicinus Scudder, 1869, and cantans Fuesslin, 1775. This STHC is compared to that proposed by Desutter-Grandcolas (2003) and is found to require a smaller amount of transformation to explain the observed morphologies. The favoured STHC implies that the stridulatory fi le is located along the same vein in all scrutinized taxa (viz. CuPb). Current phylogenetic hypotheses cannot rule out that the fi le was acquired once only. Furthermore, multiple losses explain the observed distribution more plausibly than multiple acquisitions of a complex structure. A new type of wing venation transformation is evidenced, referred to as tracheal un-capture. It involves a vein abandoning its usual course for another, and leaving a remnant of its previous course, in the form of a cross-vein-like structure (‘phantom vein’). The taxon Grylloptera is defined under cladotypic nomenclature, and is the lineage in which the character state ‘on ventral side, right and/or left forewings with a row of teeth (‘fi le’) located along CuPb’, as exhibited by viridissimus Linnaeus, 1758: p. 430 and campestris Linnaeus, 1758: p. 428, has been acquired. Type material is designated.
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Full frame distilled prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. Learned from the 10,348 direct Codex labels and 10,348 direct Gemma labels. Candidate is the union of thresholded teacher heads; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels or direct frontier model labels.
Codex and Gemma teacher scores by category
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.001 | 0.000 |
| Meta-epidemiology (narrow) | 0.000 | 0.000 |
| Meta-epidemiology (broad) | 0.000 | 0.000 |
| Bibliometrics | 0.000 | 0.000 |
| Science and technology studies | 0.000 | 0.000 |
| Scholarly communication | 0.000 | 0.000 |
| Open science | 0.000 | 0.000 |
| Research integrity | 0.000 | 0.000 |
| Insufficient payload (model declined to judge) | 0.000 | 0.000 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from it