Subspeciation within a punctuated equilibrium evolutionary event: Phylogenetic history of the Lower-Middle Ordovician<i>Paroistodus originalis—P. horridus</i>complex (Conodonta)
Bibliographic record
Abstract
A microevolutionary event involving the conodont Paroistodus lineage is documented in the Gualcamayo Formation (Middle Ordovician), Argentine Precordillera. A detailed sampling of limestones throughout the upper part of the San Juan Formation and the lower member of the Gualcamayo Formation yielded over 14,000 well-preserved conodont elements. Paroistodus originalis (Sergeeva, 1963) was recorded through the upper 230 m of the San Juan Formation and the lower member (10 m thick) of the Gualcamayo Formation. The derived species Paroistodus horridus (Barnes and Poplawski, 1973) was recorded throughout the middle member of the Gualcamayo Formation (65 m thick). The intermediate linking forms between both species are identified as two new taxa: Paroistodus horridus primus Albanesi, 1998b, and P. h. secundus Albanesi, 1998b. They were recorded in the uppermost 70 cm of the lower member. Apparently, the speciation event occurred under stressed environmental conditions with the drowning of the carbonate platform, i.e., the San Juan Formation, and the beginning of a deeper and restricted environment represented by the Gualcamayo black shales. The demise of the carbonate production was caused by a sea level rise and a significant influx of volcanic ashes. The punctuated speciation event occurred within an allopatric setting while the Precordillera occupied an isolated (Iapetus) oceanic position in its overall drift from Laurentia to Gondwana.
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How this classification was reachedexpand
Full frame distilled prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. Learned from the 10,348 direct Codex labels and 10,348 direct Gemma labels. Candidate is the union of thresholded teacher heads; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels or direct frontier model labels.
Codex and Gemma teacher scores by category
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.001 | 0.000 |
| Meta-epidemiology (narrow) | 0.000 | 0.000 |
| Meta-epidemiology (broad) | 0.001 | 0.000 |
| Bibliometrics | 0.000 | 0.000 |
| Science and technology studies | 0.000 | 0.001 |
| Scholarly communication | 0.000 | 0.000 |
| Open science | 0.001 | 0.000 |
| Research integrity | 0.000 | 0.000 |
| Insufficient payload (model declined to judge) | 0.003 | 0.000 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from itClassification
machine, unvalidatedMachine predicted; a candidate call from one teacher head, not a consensus.
How this classification was reached, model by model and score by score, is at the end of the page under "How this classification was reached".