Eureka: discovery of male Turgiditarsus Schillhammer reveals its placement in Acylophorina and resolves phylogenetic relationships within the subtribe (Coleoptera: Staphylinidae: Staphylininae)
Bibliographic record
Abstract
Turgiditarsus Schillhammer comprises some of the most bizarre and rarely encountered rove beetles in the subfamily Staphylininae. The genus is presently known from only 4 female specimens comprising three species in the eastern Oriental region. Its previous placement in Anisolinina was based primarily on superficial similarity, much before recent advances in the systematics of tribe Staphylinini. Total evidence phylogenetic analysis of a broad taxon sample within Staphylinini, including the first male specimen of Turgiditarsus, revealed that the genus should be placed in the morphologically diverse and relictual Acylophorina, a subtribe with one of the oldest crown group ages in the latest Early Cretaceous. Turgiditarsus is redescribed and two new species, T. eureka Schillhammer and Brunke sp.n. and T. vietnamensis Schillhammer and Brunke sp.n., are described, bringing the total number to five. Amacylophorus Smetana stat.n., a subgenus of Acylophorus Gravenhorst, is raised to genus rank and redescribed, with the following new combination: Amacylophorus pratensis (LeConte) comb.n. Stevensia Cameron and its single species, recently moved to this subtribe, are redescribed. All genera of Acylophorina are diagnosed and included in a novel world key. A species level key is updated for Turgiditarsus. Further systematic work within diverse Acylophorus is needed to re-assess the status of Acylohsellus, Paratolmerus and the remaining subgenera of Acylophorus.
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How this classification was reachedexpand
Full frame machine prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. The Gemma side is a direct model label for every work in the frame, read from the title-only record. The Codex side is a classifier learned from the 10,348 direct Codex labels and calibrated to design-weighted sample rates; fields without enough sample support carry no Codex call. Candidate is the union of the two sides; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels.
Distilled classifier scores by category (both heads)
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.000 | 0.000 |
| Meta-epidemiology (narrow) | 0.000 | 0.000 |
| Meta-epidemiology (broad) | 0.000 | 0.000 |
| Bibliometrics | 0.000 | 0.000 |
| Science and technology studies | 0.000 | 0.000 |
| Scholarly communication | 0.000 | 0.001 |
| Open science | 0.000 | 0.000 |
| Research integrity | 0.000 | 0.000 |
| Insufficient payload (model declined to judge) | 0.005 | 0.001 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from itClassification
machine, unvalidatedMachine predicted; a candidate call from one source (direct Gemma or distilled Codex), not a consensus.
How this classification was reached, model by model and score by score, is at the end of the page under "How this classification was reached".