Phylogenomics of microleafhoppers (Hemiptera: Cicadellidae: Typhlocybinae): morphological evolution, divergence times, and biogeography
Bibliographic record
Abstract
Abstract Phylogenomic analyses of datasets comprising 665 anchored loci and 234 species representing all recognized tribes and regional faunas of the leafhopper subfamily Typhlocybinae yielded well-resolved phylogenies for this group, largely robust to differences in analytical approach. The results support the 5 currently recognized tribes as monophyletic: Alebrini, Empoascini, Typhlocybini, Dikraneurini, and Erythroneurini, but the previously recognized tribes Zyginellini and Eupterygini are polyphyletic. A new tribe Beameranini is described based on the morphologically aberrant Neotropical genus Beamerana Young. Morphological characters traditionally used to recognize tribes are mostly stable but nearly all exhibit some homoplasy, with similar reductions and consolidations of some hind wing veins having occurred in independent lineages. Divergence time estimates indicate that the currently recognized tribes all arose during the mid- to late Cretaceous with some early splits between New and Old World lineages occur during the late Cretaceous, but most modern genera arising during the Paleogene and multiple transcontinental dispersal events also occur in the Paleogene.
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How this classification was reachedexpand
Full frame machine prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. The Gemma side is a direct model label for every work in the frame, read from the title-only record. The Codex side is a classifier learned from the 10,348 direct Codex labels and calibrated to design-weighted sample rates; fields without enough sample support carry no Codex call. Candidate is the union of the two sides; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels.
Distilled classifier scores by category (both heads)
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.000 | 0.001 |
| Meta-epidemiology (narrow) | 0.000 | 0.000 |
| Meta-epidemiology (broad) | 0.000 | 0.000 |
| Bibliometrics | 0.002 | 0.001 |
| Science and technology studies | 0.000 | 0.000 |
| Scholarly communication | 0.001 | 0.000 |
| Open science | 0.000 | 0.000 |
| Research integrity | 0.000 | 0.000 |
| Insufficient payload (model declined to judge) | 0.001 | 0.000 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from itClassification
machine, unvalidatedMachine predicted; a candidate call from one source (direct Gemma or distilled Codex), not a consensus.
How this classification was reached, model by model and score by score, is at the end of the page under "How this classification was reached".