Sensory Reconstruction of the Fossil Lorisid Mioeuoticus: Systematic and Evolutionary Implications
Bibliographic record
Abstract
The fossil record of lorises and pottos (family Lorisidae) potentially dates back to the late Oligocene of Namibia, but a later moderate diversification of this family occurred during the Miocene of Africa and Asia. In the African Miocene, the family Lorisidae is represented solely by one genus: Mioeuoticus. The phyletic position of Mioeuoticus has been a source of debate, as it has been suggested to belong to either the stem of the family Lorisidae or to be further nested within lorisids, as a sister to the African potto clade (subfamily Perodicticinae). Reconstructing the internal sensory anatomy of Mioeuoticus shipmani (KNM-RU 2052) could shed some light on this debate and possibly clarify how modern lorisoid olfactory and visual sensitivity and locomotor abilities evolved. Here, we collected data from the nasal turbinals, bony labyrinths, and orbits of Mioeuoticus shipmani from the early Miocene of Rusinga Island, Kenya. These results are consistent with Mioeuoticus, having developed typical modern lorisid behaviour (i.e., slow locomotion, nocturnal activity pattern) and olfactory abilities consistent with modern representatives. However, the arrangement of the nasal turbinals shows an intermediate state between lemuroids and lorisoids that is most consistent with a basal position of Mioeuoticus within the family Lorisidae or even the superfamily Lorisoidea.
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How this classification was reachedexpand
Full frame distilled prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. Learned from the 10,348 direct Codex labels and 10,348 direct Gemma labels. Candidate is the union of thresholded teacher heads; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels or direct frontier model labels.
Codex and Gemma teacher scores by category
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.000 | 0.000 |
| Meta-epidemiology (narrow) | 0.000 | 0.000 |
| Meta-epidemiology (broad) | 0.000 | 0.000 |
| Bibliometrics | 0.000 | 0.000 |
| Science and technology studies | 0.000 | 0.000 |
| Scholarly communication | 0.000 | 0.000 |
| Open science | 0.000 | 0.000 |
| Research integrity | 0.000 | 0.000 |
| Insufficient payload (model declined to judge) | 0.000 | 0.000 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from itClassification
machine, unvalidatedMachine predicted; a candidate call from one teacher head, not a consensus.
How this classification was reached, model by model and score by score, is at the end of the page under "How this classification was reached".