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Record W4407070057 · doi:10.1016/j.iotcps.2025.01.001

Generative AI in cybersecurity: A comprehensive review of LLM applications and vulnerabilities

2025· review· en· W4407070057 on OpenAlexaff
Mohamed Amine Ferrag, Fatima Alwahedi, Ammar Battah, Bilel Cherif, Abdechakour Mechri, Norbert Tihanyi, Tamás Bisztray, Mérouane Debbah

Bibliographic record

VenueInternet of Things and Cyber-Physical Systems · 2025
Typereview
Languageen
FieldComputer Science
TopicNetwork Security and Intrusion Detection
Canadian institutionsConcordia University
Fundersnot available
KeywordsComputer securityGenerative grammarComputer scienceArtificial intelligence

Abstract

fetched live from OpenAlex

This paper provides a comprehensive review of the future of cybersecurity through Generative AI and Large Language Models (LLMs). We explore LLM applications across various domains, including hardware design security, intrusion detection, software engineering, design verification , cyber threat intelligence , malware detection , and phishing detection . We present an overview of LLM evolution and its current state, focusing on advancements in models such as GPT-4, GPT-3.5, Mixtral-8x7B, BERT , Falcon2, and LLaMA. Our analysis extends to LLM vulnerabilities, such as prompt injection, insecure output handling, data poisoning, DDoS attacks, and adversarial instructions. We delve into mitigation strategies to protect these models, providing a comprehensive look at potential attack scenarios and prevention techniques. Furthermore, we evaluate the performance of 42 LLM models in cybersecurity knowledge and hardware security, highlighting their strengths and weaknesses. We thoroughly evaluate cybersecurity datasets for LLM training and testing, covering the lifecycle from data creation to usage and identifying gaps for future research. In addition, we review new strategies for leveraging LLMs, including techniques like Half-Quadratic Quantization (HQQ), Reinforcement Learning with Human Feedback (RLHF), Direct Preference Optimization (DPO), Quantized Low-Rank Adapters (QLoRA), and Retrieval-Augmented Generation (RAG). These insights aim to enhance real-time cybersecurity defenses and improve the sophistication of LLM applications in threat detection and response. Our paper provides a foundational understanding and strategic direction for integrating LLMs into future cybersecurity frameworks , emphasizing innovation and robust model deployment to safeguard against evolving cyber threats.

Fetched live from OpenAlex and de-inverted. Abstracts are not stored in this database: the inverted indexes are 8.6 GB of the frame’s 9.3 GB of text, and the host has 13 GB free.

How this classification was reachedexpand

Full frame machine prediction

Teacher imitation

Not calibrated prevalence, not ground truth. Human validation pending. The Gemma side is a direct model label for every work in the frame, read from the title-only record. The Codex side is a classifier learned from the 10,348 direct Codex labels and calibrated to design-weighted sample rates; fields without enough sample support carry no Codex call. Candidate is the union of the two sides; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels.

metaresearch head score (Codex)0.002
metaresearch head score (Gemma)0.007
Version: metacan-v3-hybrid-931329e0061cValidation status: machine_predicted_unvalidated
Candidate categoriesnone
Consensus categoriesnone
DomainCandidate signal: none · Consensus signal: none
Study designCandidate signal: Not applicable · Consensus signal: none
GenreCandidate signal: Review · Consensus signal: Review
Teacher disagreement score0.005
Threshold uncertainty score0.017

Distilled classifier scores by category (both heads)

CategoryCodexGemma
Metaresearch0.0020.007
Meta-epidemiology (narrow)0.0010.001
Meta-epidemiology (broad)0.0010.001
Bibliometrics0.0030.003
Science and technology studies0.0000.002
Scholarly communication0.0020.004
Open science0.0020.002
Research integrity0.0020.003
Insufficient payload (model declined to judge)0.0050.002

Machine scores (provisional)

The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.

Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.

Opus teacher head0.018
GPT teacher head0.299
Teacher spread0.280 · how far apart the two teachers sit on this one work
Validation statusscore_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from it

Classification

machine, unvalidated

Machine predicted; a candidate call from one source (direct Gemma or distilled Codex), not a consensus.

The models applied no category: nothing in the taxonomy fit this work.
Study designNot applicable
Domainnot available
GenreReview

How this classification was reached, model by model and score by score, is at the end of the page under "How this classification was reached".

Quick stats

Citations74
Published2025
Admission routes1
Has abstractyes

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