Sclerocoelus xynos Kuwahara, Marshall & Paiero, 2025, sp. nov.

Sclerocoelus xynos sp. nov. urn:lsid:zoobank.org:act: A57DAF12-DABF-4669-A254-85C1EA5AF8F7 Figs 9I, 147–149 Etymology The species name (from the Greek ‘ xynos ’, meaning ‘common’) reflects the abundance and widespread distribution of this species. Material examined Holotype VENEZUELA • ♂; Mérida, 6 km S of Azulita, near road; 3 May 1988; S.A. Marshall leg.; DEBU. Paratypes BOLIVIA – La Paz • 1 ♂, 2 ♀♀; 10 km NW of Caranavi, road to ENTEL tower; 1700 m a.s.l.; 13 Apr. 2001; S.A. Marshall leg.; bamboo dung pans; CBFC • 1 ♂; 23 km NE of Caranavi; 14 Apr. 2001; S.A. Marshall leg.; roadside/stream sweep; CBFC. ECUADOR – Napo • 11 ♂♂, 2 ♀♀; 4.2 km S of Cosanga, pipeline trail; 2200 m a.s.l.; 7 Nov. 1999; S.A. Marshall leg.; QCAZ • 5 ♂♂, 1 ♀; 5 km N of El Chaco; 15 Feb. 1983; M. Sharkey leg.; Malaise trap and wet net; CNCI • 2 ♀♀; El Chaco; 2000 m a.s.l.; 15-23 Feb. 1983; L. Masner and M. Sharkey leg.; Malaise trap; CNCI. VENEZUELA – Lara • 9 ♂♂, 1 ♀; Yacambu; 1200 m a.s.l.; 7 May 1981; H. Townes leg.; cloud forest; CNCI. – Mérida • 7 ♂♂, 3 ♀♀; same data as for holotype; DEBU • 5 ♂♂; Jají; 27 Apr. 1988; S.A. Marshall leg.; cloud forest, flight-intercept trap; DEBU • 12 ♂♂, 2 ♀♀; Jají to La Azulita; 3 May 1988; S.A. Marshall leg.; roadside sweeps; DEBU • 1 ♂; Los Chorros; 1–5 Apr. 1988; S.A. Marshall leg.; flight-intercept trap; DEBU • 2 ♂♂; same data as for preceding; leaf debris; DEBU • 5 ♂♂, 3 ♀♀; Los Chorros; 23 Apr. 1988; S.A. Marshall leg.; sweeps; DEBU • 3 ♂♂, 5 ♀♀; Mucui, 10 km E of Tobay; 2000 m a.s.l.; 28 Apr. 1981; H. Townes leg.; CNCI • 1 ♂; Santa Rosa Road; 24 Apr. 1988; S.A. Marshall leg.; vegetation along trail; DEBU • 1 ♂; Santa Rosa Trail; 23–30 Apr. 1988; S.A. Marshall leg.; flight-intercept trap; DEBU. Other material examined ARGENTINA – Salta • 7 ♂♂, 50 ♀♀; 22 km N of La Caldera, El Ucumar; 1550 m a.s.l.; 2–30 Dec. 1987; S. and J. Peck leg.; subtropical humid forest, Malaise/flight-intercept trap; CNCI • 1 ♂; 40 km N of Salta, Camino La Cornisa; 1500 m a.s.l.; 27 Feb. 1992; S.A. Marshall leg.; forest; DEBU. BOLIVIA – La Paz • 1 ♂; 10 km NW of Caranavi, road to ENTEL tower; 15°46′35″ S, 67°35′48″ W; 1400 m a.s.l.; 13 Apr. 2001; S.A. Marshall leg.; DEBU • 1 ♂, 2 ♀♀; same data as for preceding; dung pans; DEBU • 1 ♂, 2 ♀♀; same data as for preceding; 1700 m a.s.l.; bamboo dung pans; DEBU • 1 ♂; 23 km NE of Caranavi; 14 Apr. 2001; S.A. Marshall leg.; roadside/stream sweep; DEBU • 3 ♀♀; Chulumani, Apa-Apa Reserve; 16°21′15″ S, 67°30′21″ W; 2000 m a.s.l.; 1–3 Apr. 2001; S.A. Marshall leg.; pan traps; DEBU • 1 ♂; same data as for preceding; 1 Apr. 2001; sweep; DEBU. – Santa Cruz • 1 ♂; Siringalito, near Pampa Grande; 2 Oct. 1996; Bettella and Rossi leg.; DEBU • 1 ♂; Yungas de la Siberia, 26.4 km NW of Comarapa; 2640 m a.s.l.; 28 Jan. 1999; R. Anderson leg.; Yungas, mixed litter; DEBU. ECUADOR – Loja • 1 ♀; Podocarpus National Park, Bombuscaro, Higuerones trail; 1000 m a.s.l.; 21–25 Feb. 2009; M. Pollet and A. De Braekeleer leg.; yellow pans; RBINS • 1 ♀; Podocarpus National Park, Cajanuma, Bosque Nublado trail; 3000 m a.s.l.; 20–27 Feb. 2009; M. Pollet and A. De Braekeleer leg.; yellow pans; RBINS. – Napo • 1 ♀; 4.8 km W of El Chaco; 1750 m a.s.l.; 7 Nov. 1999; S.A. Marshall leg.; DEBU • 1 ♂; 17 km NE of Baeza; 1400 m a.s.l.; 3–6 Mar. 1976; S. Peck leg.; dung trap; DEBU • 30 ♂♂, 22 ♀♀; Baeza; 1500 m a.s.l.; 16–19 May 1987; L.D. Coote and B.V. Brown leg.; wet montane rainforest, Malaise trap; DEBU • 62 ♂♂, 34 ♀♀; Baeza; 1550 m a.s.l.; 15–19 May 1987; L.D. Coote and B.V. Brown leg.; wet montane rainforest/pasture, Malaise trap off stone trail; ROME • 6 ♂♂, 6 ♀♀; Baeza; 1700 m a.s.l.; 16–19 May 1987; L.D. Coote and B.V. Brown leg.; wet montane rainforest/pasture, near small creek, Malaise head; ROME • 1 ♂; Baeza; 5 Mar. 1979; S.A. Marshall leg.; DEBU • 3 ♂♂, 6 ♀♀; Baeza, W along road; 1500 m a.s.l.; 16–19 May 1987; L.D. Coote and B.V. Brown leg.; montane rainforest/pasture, Malaise trap; DEBU • 3 ♂♂, 1 ♀; El Chaco; 2000 m a.s.l.; 15–23 Feb. 1983; L. Masner and M. Sharkey leg.; Malaise trap; CNCI • 1 ♀; Quito–Baeza Pass; 4000 m a.s.l.; 4–8 Nov. 1999; S.A. Marshall leg.; pan traps in moss; DEBU • 1 ♀; SierrAzul Lodge, 14 km W of Cosanga; 2200 m a.s.l.; 5 Nov. 1999; S.A. Marshall leg.; forest, sweep; DEBU • 1 ♂, 1 ♀; SierrAzul Reserve, 14 km W of Cosanga; 0°40′55″ S, 77°56′09″ W; 2200 m a.s.l.; 9–10 May 2002; Buck and Paiero leg.; Malaise trap; DEBU • 1 ♀; same data as for preceding; 9 May 2002; M. Buck leg.; sweep trail; DEBU • 2 ♂♂, 1 ♀; same data as for preceding; 9–10 May 2002; M. Buck leg.; treefall, yellow pans; DEBU • 1 ♀; same data as for preceding; 10–11 May 2002; Marshall and Paiero leg.; debu00194781/ MYCRO919-21 sequenced for CO1–5′; DEBU • 1 ♂, 3 ♀♀; Yanayacu Biological Station; 0°36′ S, 77°53′ W; 2000–2500 m a.s.l.; 25 Apr.–13 May 2019; Yau and Marshall leg.; DEBU. – Pichincha • 2 ♂♂, 4 ♀♀; 3.5 km SE of Tandayapa; 28 Oct. 1999; S.A. Marshall leg.; in green leaf litter (treefall); DEBU • 7 ♂♂, 5 ♀♀; 7 km SE of Nanegalito, trout farm ‘ San Jose’; 1500 m a.s.l.; 27–30 Oct. 1999; S.A. Marshall leg.; riverine forest, sweeping treefalls; DEBU • 1 ♂, 4 ♀♀; Bellavista Cloud Forest Reserve; 0°01′13″ S, 78°40′30″ W; 2200 m a.s.l.; 9–13 May 2009; S.A. Marshall leg.; DEBU • 1 ♂, 1 ♀; same data as for preceding; 1 May 2011; pans near treefall DEBU • 4 ♂♂, 5 ♀♀; Bellavista Reserve; 2150 m a.s.l.; 30 Oct. 1999; S.A. Marshall leg.; on sifted litter; DEBU • 1 ♂, 1 ♀; same data as for preceding; 2200 m a.s.l.; 30 Oct. 1999; DEBU • 1 ♀; Bellavista Reserve, ridge trail; 2200 m a.s.l.; 28–30 Oct. 1999; pans near dung; S.A. Marshall leg.; DEBU • 7 ♂♂, 2 ♀♀; Bellavista Reserve, ridge trail; 28 Oct. 1999; S.A. Marshall leg.; sweeping/aspirating; DEBU • 1 ♂; Bellavista Reserve, trail B; 30 Oct. 1999; S.A. Marshall leg.; sweeping; DEBU • 1 ♀; Cotopaxi National Park, Quebrada Mishahuaycu; 3600 m a.s.l.; 16 Oct.–8 Nov. 1999; S.A. Marshall leg.; along stream, pan traps; DEBU • 1 ♀; Maquipucuna Biological Reserve, main trail; 1300 m a.s.l.; 27 Oct. 1999; S.A. Marshall leg.; DEBU • 1 ♀; Maquipucuna Biological Reserve, river trail; 1200 m a.s.l.; 29 Oct. 1999; S.A. Marshall leg.; sweep; DEBU. – Zamora-Chinchipe • 1 ♀; 8 km NW of Zamora, mouth of Río Sabanilla; 1420 m a.s.l.; 1 Nov. 1987; C. Young, R. Davidson and J. Rawlins leg.; CMNH • 2 ♀♀; San Francisco Biological Reserve, Atajo Trail; 3°58′30″ S, 79°04′25″ W; 2000 m a.s.l.; 14–18 Feb. 2009; M. Pollet and A. De Braekeleer leg.; yellow pan traps, RBINS • 4 ♂♂, 15 ♀♀; same data as for preceding; 18–25 Feb. 2009; RBINS • 1 ♂, 1 ♀; same data as for preceding; 25 Feb.–3 Mar. 2009; RBINS • 4 ♂♂, 1 ♀; San Francisco Biological Reserve, Canal Trail; 3°58′30″ S, 79°04′25″ W; 2000 m a.s.l.; 18–25 Feb. 2009; M. Pollet and A. De Braekeleer leg.; blue pan traps; RBINS • 8 ♂♂, 14 ♀♀; same data as for preceding; 13–18 Feb. 2009; red pan traps; RBINS • 1 ♂, 4 ♀♀; same data as for preceding; yellow pan traps; RBINS • 16 ♂♂, 12 ♀♀; same data as for preceding; 17–25 Feb. 2009; DEBU • 19 ♂♂, 19 ♀♀; same data as for preceding; 18–25 Feb. 2009; RBINS • 6 ♂♂, 16 ♀♀; same data as for preceding; 25 Feb.–3 Mar. 2009; DEBU • 7 ♂♂, 12 ♀♀; same data as for preceding; white pan traps; DEBU • 3 ♂♂, 5 ♀♀; San Francisco Biological Reserve, Río San Francisco Trail; 3°58′30″ S, 79°04′25″ W; 2000 m a.s.l.; 18–25 Feb. 2009; M. Pollet and A. De Braekeleer leg.; yellow pan traps; RBINS • 2 ♀♀; same data as for preceding; 25 Feb.–3 Mar. 2009; RBINS. PERU – Cusco • 3 ♂♂, 1 ♀; Cock-of-the-Rock Lodge; 13°03′21″ S, 71°32′46″ W; 1380 m a.s.l.; 18–20 Oct. 2006; J. Skevington leg.; Malaise trap; DEBU • 34 ♂♂, 45 ♀♀; Cock-of-the-Rock Lodge, NE of Paucartambo; 13°03′18″ S, 71°32′42″ W; 1120 m a.s.l.; 4–9 Nov. 2007; D. Brzoska leg.; flight-intercept trap; DEBU. – Junín • 3 ♀♀; Pampa Hermosa Lodge, 22 km N of San Ramon; 10°59′18″ S, 75°25′30″ W; 1220 m a.s.l.; 24–27 Nov. 2007; D. Brzoska leg.; flight-intercept trap; DEBU. VENEZUELA – Aragua • 1 ♂; Henri Pittier National Park, Rancho Grande; 1500 m a.s.l.; 3 Sep. 1992; L. Masner leg.; DEBU • 1 ♀; Henri Pittier National Park, Rancho Grande, La Toma trail; 9 Apr. 1994; L. Masner leg.; DEBU • 2 ♀♀; Rancho Grande, La Cumbre cloud forest; 1500 m a.s.l.; 1–10 Aug. 1987; Bordon and Peck leg.; flight-intercept trap; DEBU. – Lara • 1 ♀; Yacambú; 1200 m a.s.l.; 7 May 1981; H. Townes leg.; cloud forest; CNCI. – Mérida • 1 ♂, 1 ♀; Jají to La Azulita; 3 May 1988; S.A. Marshall leg.; roadside sweeps; DEBU • 9 ♂♂, 13 ♀♀; Mérida, Los Chorros; 2300 m a.s.l.; 23–30 Apr. 1988; S.A. Marshall leg.; flight-intercept trap; DEBU • 1 ♀; Mérida, Los Chorros; 1–5 May 1988; S.A. Marshall leg.; dung trap; DEBU • 1 ♂, 1 ♀; same data as for preceding; leaf litter, flight-intercept trap; DEBU • 1 ♀; Mérida, Los Chorros, Santa Rosa Road; 30 Apr. 1988; S.A. Marshall leg.; sweep; DEBU • 1 ♂, 1 ♀; Mérida, Los Chorros; 23 Apr. 1988; S.A. Marshall leg.; sweep; DEBU • 1 ♂, 1 ♀; Mérida, Santa Rosa Road; 25–30 Apr. 1988; S.A. Marshall leg.; pasture/cloud forest edge, Malaise trap; DEBU • 9 ♂♂, 11 ♀♀; Santa Rosa Trail; 1500 m a.s.l.; 24–30 Apr. 1988; S.A. Marshall leg.; flight-intercept trap along spring; DEBU. – Táchira • 1 ♀; 38 km NE of San Cristóbal; 2100 m a.s.l.; 18–20 Jun. 1974; S. Peck leg.; carrion; DEBU. – Trujillo • 2 ♂♂, 1 ♀; 10 km E of Bocono, Laguna de Lucerdo; 1760 m a.s.l.; 3 Mar. 1995; S.A. Marshall leg.; slash/compost; DEBU • 1 ♀; Bocono–Guaramacal Road; 2130 m a.s.l.; S.A. Marshall; sweep wet litter; DEBU • Guaramacal National Park; 2000–3000 m a.s.l.; 26 Aug.–1 Sep. 1992; L. Masner leg.; car net; DEBU. Description BODY (Fig. 147A). Length 2.3–3.8 mm. Head dark brown, very bottom edge of frons orange; face and gena dark brown, antennae orange; frontal vitta, interfrontal plate and orbital plate slightly paler and shiny. Frontal width 2.3–2.4 × frontal height. Two pairs of strong interfrontal bristles surmounting a very fine third pair; anterior orbital 0.4–0.5× length of posterior. Palpus yellow. Ey

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