Bromus lanatipes Rydberg 1906

15. Bromus lanatipes (Shear) Rydberg (1906: 52). Fig. 50, 51. Basionym: Bromus porteri var. lanatipes Shear (1900: 37). Bromus anomalus var. lanatipes (Shear) Hitchcock (1933: 449). Bromopsis lanatipes (Shear) Holub (1973: 168). Type:— UNITED STATES OF AMERICA. Colorado: Idaho Springs, 27 August 1895, C.L. Shear 739 (holotype US-81588!, isotype RM-150554!). Plants perennial, not rhizomatous. Culms to 120 cm tall, (1–) 2–3 mm wide at base, glabrous or pubescent below inflorescences; nodes 3–4, light to dark brown, pubescent. Leaf sheaths lanate, occasionally densely pilose, hairs up to 1.2 mm long; auricles absent; ligules 0.5–1.5 mm long, glabrous; blades up to 22 cm × 4–7 mm, flat, adaxial and abaxial surfaces glabrous or minutely pubescent, margins serrulate. Panicles 16–20 cm × 2–8 cm, open, nodding, branches ascending to spreading, shorter or longer than spikelets, scabrous, 1–4 spikelets per branch. Spikelets 1.7–3 cm long, florets 7–9-flowered, elliptic to lanceolate, terete to moderately laterally compressed; glumes glabrous, pubescent along margins or pubescent throughout, margins hyaline, midnerves glabrous or scabrous; lower glumes 4–6 mm long, lanceolate, 1-nerved, green along the nerve, apices acute; upper glumes 6.5–8 mm long, obovate-lanceolate, 3-nerved, green along and between the nerves, apices acute or mucronate, mucros up to 1 mm long; lemmas 8–10 mm × 1–2 mm, elliptic to lanceolate, rounded over the backs, apices truncate, 5-7-nerved, green along and between the nerves, densely pubescent or hairs restricted to margins, marginal hairs sometimes longer than those on back, hairs up to 0.6 mm long; awns 2–6.5 mm long, arising 0–0.5 mm below lemma apex, straight; paleas usually equal in length to the lemma, 7–10 mm long, backs pubescent, keels ciliate, cilia up to 0.2 mm long; anthers 2–4 mm long; caryopses 6−7 mm long, light brown. 2 n = 28 (Wagnon 1952, Ward & Spellenberg 1988). Distribution: ―Native. Bromus lanatipes is known only from northern Coahuila in México, near the United States border (Fig. 52). In the United States its range includes western Texas, New México, Arizona, Nevada, Utah, Colarado and Wyoming (Pavlick & Anderton 2007). Ecology: — Bromus lanatipes is found along ridgetops and steep slopes; associated with Pinus spp., Abies, Pseudotsuga menziesii, Quercus spp., Juniperus, Cupressus arizonica, Arbutus, Salvia, Salix Linnaeus (1753: 1753), Rhamnus and Ceanothus. Elevation: 1676–2627 m. Common Names: ―Woolly brome, shaggy brome (English); bromo veluludo (Spanish). Comments: ― Bromus lanatipes is distinguished by its lanate lower sheaths. Aphotograph of this character is given in Allred (1993: 334). Circumscribing this taxon required examination of material from the adjacent United States, which is cited below. Specimens Examined:― MÉXICO. Coahuila: ca. 12 (air) mi Eof Boquillas in Sierra del Carmen, 10.4 (rd) mi NW of Rancho El Jardín, 29.2°N, 102.23°W, 5650 ft [1722 m], J. Henrickson 11512 (RSA-POM-664819); ca. 12 (air) mi Eof Boquillas, 8.3 (rd) mi NW of Rancho El Jardín, 29.1667°N, 102.23°W, 5500 ft [1676 m], 27 July 1973, J. Henrickson 11487 (RSA-POM-665612); Maderas del Carmen, 12.2 mi NW of Pilares near "old cabin", 28.9401°N, 102.5978°W, 2300 m, 7 September 2005, P.M. Peterson & J. Valdés-Reyna 18889 (ANSM, CAN, MO, US); Maderas del Carmen, 16.3 mi NE of Los Pilares on road towards Campo Dos, 28.9626°N, 102.5643°W, 2235 m, 21 September 2007, P.M. Peterson, J.M. Saarela, S. Lara Contreras & J. Reyna Álvarez 20983 (CAN, MO, P-03216868, US); Maderas del Carmen, 17.7 mi NE of Los Pilares on road towards Campo Dos, 28.9645°N, 102.56°W, 1954 m, 21 September 2007, P.M. Peterson, J.M. Saarela, S. Lara Contreras & J. Reyna Álvarez 21010 (CAN, MO, US); Sierra El Jardín, 29.0832°N, 102.6371°W, 2100 m, 3 September 2006, P.M. Peterson & S. Lara-Contreras 19938 (CAN, MO, US); Madera del Carmen, at Campo Dos, 29.0106°N, 102.6083°W, 2627 m, 16 September 2012, P.M. Peterson & K. Romaschenko 24531 (US). UNITED STATES OF AMERICA. Arizona: Navajo Indian Reservation, about the Nend of the Carizo Mtns., 29 July 1911, P.C. Standley 7386 (US-686353); Cococino Co., 30 mi Eof Canyon Padre, Flagstaff, 7–11 August 1915, A.S. Hitchcock 13243 (US-906049); Cochise Co., Huachuca Mtns., September 1893, J.G. Lemmon s.n. (US, 2 sheets); Dos Cabezos, 31 October 1937, L.N. Goodding s.n. (US-1721928). New México: Gray’s Peak, 6500 ft, F.S. Earle 162 (US-1008675); Grant Co., about the S end of the Black Range, 6000 ft, 1904, O.B. Metcalfe s.n. (US-690339); Santa Fe Co., Glorieta, 2 July 1926, A.S. Hitchcock 22994 (US-1296512); Glorieta, 24 August 1910, E.O. Wooton s.n. (US-726304); Santa Fe, 3 July 1926, A.S. Hitchcock 23011 (US-1296516); Santa Fe, 1884, G. Vasey s.n. (US-1008802); Lincoln Co., Gray, 6000–6500 ft, July 1900, F.S. Earle & E.S. Earle 162 (US-382342); Ft. Stanton on El Paso road, 3 August 1937, L.N. Goodding 3514 (US-1723783); between Ft. Stanton and Ruidoso, 26 August 1937, L.N. Gooding & C. Goodding 3519 (US-1723785); San Miguel Co., Las Vegas, 23 June 1891, L.H. D. s.n. (US-749847); vicinity of Las Vegas, 27 June 1928, G.E. Osterhout 7048 (US-1389144). Oklahoma: Cimarron Co., Mesa de Maya (Black Mesa), 3 mi Nof Kenton, 9 July 1947, C.M. Rogers 4747 (US-2010965); Nslopes of Black Mesa, 3 mi Nof Kenton, 30 May 1952, U.T. Waterfall 10749 (US-2077766). Texas: Guadalupe Mts., 1881, [?] s.n. (US-132953); Brewster Co., Chisos Mtns., Green Gulche, below Casa Grande, 26 August 1944, C.L. Lundell 13222 (US-1913218); Jeff Davis Co., Sawtooth Mountain, Davis Mountains, 3 October 1926, E.J. Palmer 31833 (US-1297432); Trans-Peco Texas, Davis Mtns., High Mt. Livermore, 22 July 1936, L.C. Hinckley 607a (US-1871685); Uvalde Co., Black Mt., 13 August 1926, B.C. Tharp 4105 (US-1296999).

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