Quedius molochinus

Quedius molochinus (Gravenhorst, 1806) (Figs 1, 4, 9, 8C,D, 12, 13D, 20) Staphylinus molochinus Gravenhorst, 1806: 56 [Type locality:Brunsviga] Quedius denudatus Stephens, 1832:216 [Type locality: London; Suffolk] Quedius lathburii Stephens, 1832: 218 [Type locality: London; Suffolk, Orwell] Quedius niger Sahlberg, 1876: 229 [Type locality: Muonioniska] Quedius maculicornis Mulsant & Rey, 1876: 694 [Type locality: les montagnes du Lyonnais, à la Grande Chartreuse, dans les Pyrénées] Raphirus picipennis Stephens, 1833: 243 [Type locality: London] Staphylinus laevicollis Runde, 1835: 6 [Type locality: Halae] Staphylinus lapponicus Zetterstedt, 1838: 61 [Type locality: Lapponia Umensis, Lycksele; Wilhelmina; Gaskelougt] References. GඒඅඅൾඇHൺඅ (1810):302, (1827):477 (characters);SൺHඅൻൾඋG (1830): 314 (characters); (1876): 24 (distribution); MൺඇඇൾඋHൾංආ (1830): 25, (1831): 439 (distribution); (1843): 232 (characters); Rඎඇൽൾ (1835): 5 (characters); Lൺർඈඋൽൺංඋൾ (1835): 377 (characters); Cඎඋඍංඌ (1837): pl. 638 (characters); Nඈඋൽආൺඇඇ (1837): 76 (distribution); EඋංർHඌඈඇ (1839): 489, (1840): 535 (characters); Hൾൾඋ (1839): 276 (characters); SඍൾඉHൾඇඌ (1839):390 (characters);Lൺඉඈඋඍൾ (1840):177 (characters);Kංൾඌൾඇඐൾඍඍൾඋ (1848): 52 (characters); (1851): 419 (distribution); GඋൺඏൾඇHඈඋඌඍ (1847): 227 (characters); RൾൽඍൾඇൻൺർHൾඋ (1849): 709, (1857): 201, (1874): 198 (characters); Hൺඋൽඒ (1851): 32 (biology); Fൺංඋආൺංඋൾ & Lൺൻඈඎඅൻජඇൾ (1856): 537 (characters), Kඋൺൺඍඓ (1857): 500 (characters); WൺඍൾඋHඈඎඌൾ (1858): 23 (synonymy); THඈආඌඈඇ (1859): 25 (notes on types); (1860): 173 (synonymy); GൾආආංඇGൾඋ & Hൺඋඈඅൽ (1868): 570 (synonymy); Fൺඎඏൾඅ (1874): 290 (characters and distribution); Sൾංൽඅංඍඓ (1875): 267 (characters); Mඎඅඌൺඇඍ & Rൾඒ (1876): 692 (characters); Fඈඐඅൾඋ (1888): 237 (characters); Rൾංඍඍൾඋ (1909): 112 (characters); JඈHൺඇඌൾඇ (1914): 364 (characters); Gඋංൽൾඅඅං (1924): 81, (1962a): 320 (characters); Pඈඋඍൾඏංඇ (1929): 341 (characters); MඣඊඎංGඇඈඇ (1937): 3 (synonymy); Hൺඇඌൾඇ (1952): 142 (characters); Sආൾඍൺඇൺ (1958): 364 (characters and biology); (1962a): 136, (1965a): 38 (characters); (1962b): 134 (larval characters); (1964): 80, (1976a): 172, (1978a): 85, (1981): 638, (1990): 98, (1993): 50 (distribution); (1971): 133 (distribution; introduction to Nearctic); (1973): 1427 (introduction to Nearctic);KඈඋGൾ (1960b): 53 (characters); Cඈංൿൿൺංඍ (1961): 52, (1963): 423, (1978):194 (characters); Pൺඅආ (1963a):141 (characters); LඈHඌൾ (1964): 212 (characters); Dඏඈřák 1965: 89 (distribution); Pඈඍඈඍඌĸൺංൺ (1967): 82 (larval characters) Kൺඌඎඅൾ (1968a): 66 (biology); (1970): 61 (larval characters); Sඓඎඃൾർĸං (1968): 736 (distribution); Oඌൾඅ-අൺ & Zൺඇൾඍඍං (1975): 120 (biology); Bඈඋൽඈඇං (1976a): 95 (characters); SHංඅඈඏ (1975): 376 (distribution); Pඈඉൾ (1977): 31 (distribution); TඬඍH (1984):120 (characters);SൾGൾඋඌ(1987):182 (morphology);WൾඅർH (1993): 229 (morphology);Cංർൾඋඈඇං & Zൺඇൾඍඍං (1995):32 (distribution); Sඍൺඇංൾർ (1996):117 (pupal characters);Dඈඐඇංൾ &Aඋඇൾඍඍ (1996):391 (characters); Oඐൾඇ (2000): 252 (biology); AඌඌංඇG (2001): 75 (biology); UHඅංG et al. (2006): 55 (distribution); Mൺඃĸൺ (2007): 949 (distribution); Dൺඉĸඎඌ & Tൺආඎඍංඌ (2008): 33 (distribution); Cඎඉඉൾඇ et al. (2011): 15 (distribution); Sൾආංඈඇൾඇĸඈඏ et al. (2015): 335 (distribution). Type material examined. Staphylinus molochinus: Sඒඇඍඒඉൾ J (ZMHB), labelled:“6230 / Braunsberg Dahl leg./ molochinus gr.gyll. / (Para)? typus Grav. / det. H. Korge Quedius molochinus Grav. Typus!”. Quedius niger: Sඒඇඍඒඉൾ ♀ (MZH), labelled:“Tjudi / J. Sahlbg./ 1617. / Mus.Zool.H:fors Spec.typ. No 157 Qued.molochinus var.niger J. Sbg. / Syntype / http://id.luomus.fi/GZ.83 / Photographed 2020 Pekka Malinen”. Additional material examined. AUSTRIA: Arbesbach, [48.49, 14.95], 13.VII.1957, leg. Schubert (1 ♀ NMW); Admont, [47.56, 14.45], leg. H. Franz (1 ♀ NMW); Prater, [48.21, 16.39], 19.IX.1964 (1 J NHMD). CZECH REPUBLIC: Iser [Jizera] River, n. Benešov u Semil, [50.60, 15.36], 350 m, 24.VII.1982, leg.Hieke (1♀ ZMHB);Pastviny n.Žamberk, [50.09, 16.56], 31.VII.-3.VIII.1982, leg. Hieke (1 ♀ ZMHB); Podyji Nat. Park,forest, at river Dyje valley ca. 2.5 km NW of Havraníky,48.83,15.98, 240 m, sifted flood debris, 08.VI.2016, leg. A. Solodovnikov, J. Jenkins Shaw,&A. Hansen (1J NHMD); Silesia Altvater [Praděd], [50.08, 17.23], leg. Letzner (1 SDEI); Teplice nad Metují, [50.59, 16.16], 3.-5.VIII.1992, leg.Hieke (1♀ ZMHB);Vysoká Hola, NW Karlov, Malá Morávka, [50.08, 17.23], sifting litter of Pinus mugo, 25.VII.1993, leg. Zerche (1 SDEI). DENMARK: Hjelm Hede, [56.49, 8.91], 20.VII.1951, leg. J. Petersen (2 ♀♀ NHMD). BඈඋඇHඈඅආ: Øle Å, 19.IV.1965, [55.05, 15.02], leg. F. Bangsholt (1 ♀ NHMD). Fඎඇൾඇ: Bjergsted, [55.66, 11.35], 1.VI.1981, leg. M.Hansen (1 J NHMD);Ristinge Klint, [54.82, 10.61], 14.VII.1978, leg. M. Hansen (2 JJ NHMD). Zൾൺඅൺඇൽ: Furesøstien, [55.81, 12.41], 27.IX.1954, leg. J. Petersen (3 ♀♀ NHMD); Holmegårds Mose, [55.29, 11.82], 26.VI.1975, leg. M. Hansen (1 J NHMD); Ormø, [55.21, 11.45], 13.IX.1997, leg. O. Martin (1 ♀ NHMD); Rude Skov, [55.83, 12.47], 19.VIII.1948, leg. J. Petersen (1 ♀ NHMD). FINLAND: Kilpisjärvi, [69.04,20.81], 21.-22.VII.1985, leg.Hieke (1♀ ZMHB);Kiuruvesi,[63.65, 26.62], 6.VII.1951, leg. Linnavuori (1 J 2 ♀♀ ZMUO); Mutenia, Sodankylä,[68.02, 27.41], 13.IX.2001, leg.J.Itämies, O. Nenonen (1♀ ZMUO); Rymättylä,[60.37, 21.94],leg.Linnavuori (1♀ ZMUO);Saana,Enontekiö, [69.04,20.85], 10.VII.2013, M.Pentinsaari (1J ZMUO). FRANCE: Chigny,Oise River, [49.91, 3.77],Selchert (1J ZMHB). GERMANY: Bayern, Schleissheim,[48.24, 11.59], 15.V.1931, leg. Ihssen (1J ZMHB); Berlin--Marzahn, Hellersdorfer Berg, [52.53,13.58], 8.VIII.1997, leg. Hieke (1J ZMHB); Brunn, Dosse [52.91, 12.51], 16.III.1980 leg. E. Rössner (2 JJ SDEI); Engolling,Auerbach, [48.79,13.10], 5.IX.1978, leg.K.Koufrauen (1 J 1 ♀ ZMHB); Erzgebirge, Flohtelberg, [50.42, 12.95], 2.VI.1925, leg Uhmann (1 J SDEI); Finkenkrug nr. Berlin, [52.56, 13.03], 12.IV.1898, leg. Dahl (1 ♀ ZMHB); Flessenow, Schwerin, [53.75, 11.49], 4.VII.1968, leg.Uhlig (1 ZMHB); Hainleite, SW Günseroda and Seega,[51.32, 11.02], 10.VIII.1993, leg. F. Hieke (1 J ZMHB); Marburg in Hess, [50.81, 8.77], 16.IV.1904, leg. Strand (1 J ZMHB); Neuendorf, Teupitz, [52.11, 13.58], 30.IV.1992, leg. Renner (1 J ZMHB); Sarnow n.Anklam, [53.75, 13.63], 29.IX.1964, leg. F. Hieke (1 J ZMHB); Strausberg, 23.VII.1981, [52.56, 13.88], leg. H. Wendt (1 ♀ ZMHB); Thüringer Wald nr. Scheibe, [50.48, 11.05], under stones, 8.I.1957, leg. Dieckmann (1 SDEI); Willersdorf, [51.02, 8.85], VII (1 ♀ ZMHB). ITALY: Gello, [43.96, 10.45], VI.1922, leg. A. Strand (1 J NHMD); Zeccone n. Pavia, [45.26, 9.19], 17.X.1977, leg. P. Kanaar (1 J ZMHB). NORWAY: Hemnesberget, [66.22, 13.61], 8.-14.VII.1903, leg. E. Strand (2 JJ ZMHB); Kongsvinger Eis, [60.19, 12.01], 27.X.1980, leg. F. Midtgaard (1 J NHMD); Pasvik Lake, Sydvaranger, [69.34, 29.53], VI.1994, leg. G. Nietsch (1 J NMW); Utgardsjön, Rønningen,Austmarka, [60.01, 12.37], 28.VI.-10.VII.1975, leg.P.Kanaar (1 ♀ ZMHB). POLAND: Misdroy, [53.91, 14.44] (1 J ZMHB); Pölitz [Police], Pommern,[53.55, 14.57], 13.VI.1920,leg.E. Hanau (1♀ ZMHB); Riesengebirge [Krkonoše], [50.79, 15.57], 750m, moss, 16.X.1902, leg. Dahl (1 J ZMHB); Tamsel, [52.62, 14.70], 20.V.1920, leg. Kuntzen (1 ♀ ZMHB). RUSSIA: CH ංඍൺ Oൻඅൺඌඍ: Malkhanskiy Khrebet, pass to Maleta, 20 km S Maloarkhangel’sk, 50.2352 108.916, 19.-25.VIII.1998, leg. A. Shavrin (NHMD). KHൺඇඍඒ- Mൺඇඌං Aඎඍඈඇඈආඈඎඌ Dංඌඍඋංർඍ: Sovetsky region, Malaya Sosva Nature Reserve, near Cordon Hangokurt, road to Belaya Gora, 61.9504, 64.2489, 81 m, nr a roadside puddle with Polytrichum sp. on clay, 28.VI.2017, leg. A.B. Ryvkin (1 J cRyv). Kඈආං Rൾඉ: Ukhta surrounding,[63.57, 53.71], 29.VI.1974, leg.V.Shulov (6JJ 6 ♀♀ ZMHB). KඋൺඌඇඈඒൺඋඌK Kඋൺං: Niznaya Lebedyanka river, [61.96, 89.45], 23.VI.1992, leg. V.S. (1 J cRyv); Turukhanskiy Distr., Mirnoye, [62.28, 89.03] 6.-17.VII.1990, leg. L.B. Rybalov (1 J 1 ♀ cRyv). Lൾඇංඇ-G උൺൽ Oൻඅൺඌඍ: Saint-Petersburg, Sestroretsk, Rzhavaya kanava, [60.12, 29.96], 16.VIII.2009, leg.V.N.Prasolov (1J ZIN); Stary Petergof, [59.87, 29.87], forest, 7.VI.1997, in leaf litter, leg.A. Solodovnikov (1J NHMD) Yaschera, [58.92, 29.98], 18.V.1996, leg.L.Anisutkin (1J NHMD). Mඈ-ඌർඈඐ Oൻඅൺඌඍ: Moscow,Izmajlovskij Park,[55.75,37.62], 7.VII.1983, leg. A.Isaev (1J NHMD); Odintsovsky Distr., Zvenigorod Biological Station of MSU, [55.70, 36.72], mixed forest, 22.VIII.1984, leg. K.G. Mikhailov (2 JJ cRyv). MඎඋආൺඇඌK Oൻඅൺඌඍ: Kyndomys [Kindo] Peninsula,[66.65, 33.22], forest floor, 16.VII.1992, leg. Wegner (1 J ZMHB). Pൾඋආ Kඋൺං: Perm city, [58.01, 56.22], pitfall, 29.VIII.1978, leg. S.L. Esunin (1 J NHMD). Rൾඉ. ඈൿ Bඎඋඒൺඍංൺ: Barguzinsky Distr., Barguzin valley, 20 km E Ust-Barguzin, Gusikha, [53.42, 109.42], aspen forest, pitfalls, leg. V. Shilenkov (1 J NHMD); Vitimsky Nat. Preserve, lake Oron, mouth or Labaznyj creek, Oron hut, [54.82, 112.26], 27-30.VII.2000, leg. A. Shavrin (2 JJ NHMD). Rඒൺඓൺඇ Oൻඅൺඌඍ: Ryazan Prov., Spassky Distr., Oka nature reserve, [54.71, 40.85], leaf litter, 26.VI.1981, leg. Kormylutsyn & Eskov (1J 1♀ cRyv). SඏൾඋൽඅඈඏඌK Oൻඅൺඌඍ: Ural,Visim Nature Reserve, [57.37, 59.77], 25.I.1985, leg L.P. Titova (2 JJ cRyv). Tඒඎආൾඇ Oൻඅൺඌඍ: Surgutsky Distr., Yugansky Nature Reserve, Ai-Magromsy River Basin, near ‘Medvezhii Ugol’, [59.59, 74.71], 22.IX.2002, leg.A.B. Ryvkin (1 J cRyv). UඅඒൺඇඈඏඌKൺඃൺ Oൻඅൺඌඍ: Inzenskij Distr., Argash village, [53.96, 46.25]‚ birch forest, 9.VII.1995, leg.A.Isaev (2JJ NHMD). SLOVAKIA: Inovec, [48.77, 18.04], IX.1931, leg. Čepelák (1 J ZMHB). SWEDEN: Dagsås-trakten, Hall, [57.06, 12.48], 26.X.1953, leg. J.T. Skovgaard (2 JJ 3 ♀♀ NHMD); Södermanand, Ornö, [59.05, 18.41], 15.V.1946, leg. J.T. Skovgaard (1 J 1 ♀ NHMD); Stockholm, Djurgården 23.III.1945, leg. J.T. Skovgaard (1 ♀ NHMD); Vitamölle, [55.69, 14.21], 1.VIII.1926, leg.Stephensen (1J NHMD). Gඈඍඅൺඇൽ: Lau,[57.28, 18.61], 25.VI.1950 leg. J.T. Skovgaard (1 J NHMD). SWITZERLAND: Reussdelta, Seedorf, [46.89, 8.61], 435 m, VIII.1998, leg. Herger & Rezb. (1 J ZMHB). UNIT

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