Diploneis kilhamiana Jovanovska & Wilson & Hamilton & Stone 2023, sp. nov.
Bibliographic record
Abstract
Diploneis kilhamiana sp. nov. (LM Figs 349–372, SEM Figs 373–383) Valves are weakly asymmetric, linear-elliptic with parallel to weakly convex margins and round apices (Figs 349–373). Valve length is 22.5–43 μm and width is 11.5–15 μm. The axial area is linear to lanceolate, widening at the center to form a small lanceolate central area (Figs 351, 373, 376), 2.5–3.5 μm wide. Externally, the canal is linear to lanceolate, slightly expanded in the middle of the valve with two rows of cribrate (8–15 poroids) areolae narrowing into one at the valve apices (Figs 351, 373, 375). Internally, a thick non-porous slightly raised silica plate encloses the longitudinal canal (Figs 379, 381). Externally, the raphe is filiform, curved with expanded proximal ends weakly deflected to one side; the proximal ends are positioned within an elongated teardrop depression (Figs 373, 375, 376). The distal raphe ends are unilaterally bent to the same side and terminate as expanded pores at the valve face margin (Figs 374, 375, 377). Internally, the raphe is curved with simple proximal and distal ends that are slightly elevated in a depression formed by the longitudinal canal (Figs 380, 381, 383). The striae are parallel at mid-valve becoming radiate towards the apices, 9–11 in 10 μm. Striae are biseriate throughout (Figs 378, 379, 382). The striae are composed of round to rectangular areolae covered externally with fine pored cribra (15–20 poroids), 20 in 10 μm. The inter-areolar thickenings have fin-like silica ridges serrated with ca. 8–10 notched edges (Fig. 374). The fin-like silica ornamentations along the canal are slightly bent into a semi-circular shape, positioned towards the striae whereas those of the striae are only slightly bent towards the canal and change direction only at the valve mantle (Figs 374–376). The areolae increase in size towards the valve margins (Fig. 378). Internally, the alveoli open via a single elongated opening covered with a thin silica layer (Figs 380–382). The valvocopula has serrated advalvar edges (Figs 379–382). Type:— REPUBLIC OF ZAMBIA, Lake Tanganyika, Kalambo Falls Lodge, at 770 m elevation; mud, 18 m water depth, collected SCUBA diving, 8°37’25.6” S 31°11’59.7” E, H. Büscher, 1 st September 2018 (holotype designated here, circled specimen BM-108985! = Fig. 370, isotypes ANSP-GC17214!, CANA-129335!). Type material CANA-129315. Registration: http://phycobank.org/103710 Pictures of the isolated specimen:— LM micrograph on 1000× magnification (Fig. S3v). Sequence data:— Plastid gene rbc L sequence (GenBank accession: OQ 660280). Etymology:— The specific epithet ‘ kilhamiana ’ honors Dr. Susan S. Kilham, who made many contributions to the field of diatom ecology, including research in Lake Tanganyika. Ecology and distribution:— This species has been observed in Lake Tanganyika, where the water is predominantly alkaline, moderately mineral-rich and very transparent. The species is widespread in the southern, central, and northern sub-basins on the Tanzanian and Zambian sides (including Burundi, see Cocquyt 1999, fig. 16), especially at Kalambo Falls Lodge, Cape Nangu in Kasaba Bay, Mahale National Park, Kiganza Bay, as well as Isanga Bay, Chituta Bay, and the Rukoma area (see Fig. 1c–f). Diploneis kilhamiana sp. nov. inhabits sandy and muddy substrates (sometimes with mollusk shells) between 10 and 36 m water depth. A few specimens were found in 70 m water depth in the cleaned samples from Chituta Bay, and live specimens were found on the submerged rocks in the coastal areas at Jakobsen Beach near Kigoma, probably due to sediment distribution caused by currents and water turbulence at both sites. This species normally coexists with D. cocquytiana sp. nov., D. gigantea sp. nov., D. salzburgeri sp. nov., D. tenera sp. nov., D. nana sp. nov., D. fossa sp. nov., D. angusta sp. nov., and D. cristata sp. nov. Main differential characters:— Valve shape, striae pattern, and external thick fin-like ornamentations across the valve; fins more randomly distributed across the canals. Similar species:— Diploneis cocquytiana sp. nov., Diploneis heteromorphiforma Metzeltin, Lange-Bertalot & Nergui (2009: 35), and Diploneis pseudovalis Hustedt (1930: 253).
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How this classification was reachedexpand
Full frame machine prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. The Gemma side is a direct model label for every work in the frame, read from the title-only record. The Codex side is a classifier learned from the 10,348 direct Codex labels and calibrated to design-weighted sample rates; fields without enough sample support carry no Codex call. Candidate is the union of the two sides; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels.
Distilled classifier scores by category (both heads)
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.000 | 0.000 |
| Meta-epidemiology (narrow) | 0.001 | 0.000 |
| Meta-epidemiology (broad) | 0.000 | 0.000 |
| Bibliometrics | 0.001 | 0.001 |
| Science and technology studies | 0.001 | 0.000 |
| Scholarly communication | 0.001 | 0.001 |
| Open science | 0.000 | 0.001 |
| Research integrity | 0.000 | 0.000 |
| Insufficient payload (model declined to judge) | 0.006 | 0.003 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from itClassification
machine, unvalidatedMachine predicted; a candidate call from one source (direct Gemma or distilled Codex), not a consensus.
How this classification was reached, model by model and score by score, is at the end of the page under "How this classification was reached".