Calofulcinia oxynota La Greca. A. Lateral 1969

Calofulcinia oxynota La Greca Figures 4–7, 11A Calofulcinia oxynota La Greca, M. 1969, Su alcuni Mantodei dell’Australia e delle Filippine. Annali del Museo civico di storia naturale Giacomo Doria, vol. 77, pp. 633–639 [633]. Type material. The holotype female could not be located by Balderson (Balderson 1984); it is not in La Greca’s collection, nor is it in the Natural History Museum, London. La Greca’s (1969) detailed description and figures leave no doubt as to the identity of the species, however. He gives the following details for the type specimen: “ QUEENSLAND: Cairns, Barrow River, 12.IX.1966, 1 ♀ (A. Walford-Huggins leg.) (tipo).” “Barrow River” is almost certainly misspelt and refers to the Barron River in Cairns. Specimens examined. QLD: 1♁ Stewart Ck., 4km NNE Mt Spurgeon (Camp 1) 1250–1300m, 16°24’S 145°13’E, 15–20 OCT 1991, Monteith, Janetzki, Cook & Roberts (QM); 1♁ The Bluff, 11km W of Mossman 800– 1300m, 2 NOV 1983, Monteith, Yeates & Thompson (QM); 1♁ Mossman Bluff Track 10km W Mossman 1200m, 17 DEC 1988, Monteith & Thompson, pyrethrum/trees & logs (QM); 1♁ Mossman Bluff Track, 5–10km W. Mossman, Site 8 1180m, 20 DEC 1989 – 15 JAN 1990, Monteith, Thompson & ANZSES, flt. Intercept (QM); 1♁ Mossman Bluff Summit 10km W Mossman 1300m, 21 DEC 1989, Monteith, Thompson & ANZSES (QM); 1♁ 2km ESE Mossman Bluff, 9km W Mossman 1000m, 17–18 DEC 1988, Monteith & Thompson (QM); 1♁ Mt Lewis 500m past swamp, 29 DEC 1989, J. Hasenpusch (MV); 1♁ Mt Lewis, 23 DEC 1995, J. Hasenpusch (ANIC); 1♀ Bakers Blue Mt 17 km W Mt Molloy 900m, 11 SEP 1981, G. Monteith & D. Cook (QM); 1♁ same locality and collectors, 1100m, 12 SEP 1981 (QM); 1♀ same locality, 800–1000m, 30 DEC 1989 – 9 JAN 1990, ANZES Expedition (QM); 1♀ Mt Formartine South, 10km N. Kuranda 700m, 23 NOV 1990, Monteith & Thompson, pyrethrum/trees & logs (QM); 1♁ Kuranda (335m) (Top of the Range) 19 Butler Dr 16.48’S 145.38’E. (GPS), 16–31 OCT 2008, DCF Rentz, ANIC Database No. 11 000022 (ANIC); 2♁♁ same locality and collector, 18–30 SEP 2009, ANIC Database No. 11 000023 (ANIC); 1♀ same locality and collector, 1–15 OCT 2009 (ANIC); 1♁ same locality and collector, 16–31 JAN 2011, ANIC Database No. 11 000020 (ANIC); 1♁ same locality and collector, 16–31 MAR 2011, ANIC Database No. 11 000021 (ANIC); 1♁ same locality and collector, 1–15 FEB 2011, ANIC Database No. 11 000032 (ANIC); 1♁ same locality and collector, 1–15 AUG 2016 (ANIC); 1♁ same locality and collector, 16–30 SEP 2016 (ANIC); 2♁♁ same locality and collector, 1–15 NOV 2016 (ANIC); 1♁ same locality and collector, 16–31 JAN 2017 (ANIC); 1♁ same locality and collector, 16–30 APR 2017 (ANIC); 1♁ same locality and collector, 1–15 JUN 2017 (ANIC); 1♁ same locality and collector, 16–30 NOV 2017 (ANIC); 1♁ same locality and collector, 1–15 JAN 2018 (ANIC); 1♁ same locality and collector, 1–15 FEB 2018 (ANIC); 2♁♁ same locality and collector, 1–15 MAR 2018 (ANIC); 1♁ same locality and collector, 1–15 APR 2018 (ANIC); 1♁ same locality and collector, 1–15 JUL 2018 (ANIC); 1♁ same locality and collector, 16–31 OCT 2018 (ANIC); 2♁♁ same locality and collector, 16–30 NOV 2018 (ANIC); 1♁ same locality and collector, 1–15 JAN 2019 (ANIC); 1♁ same locality and collector, 16–31 JAN 2019 (ANIC); 1♁ same locality and collector, 1–15 FEB 2019 (ANIC); 1♁ same locality and collector, 16–28 FEB 2019 (ANIC); 1♁ same locality and collector, 1–15 SEP 2019 (ANIC); 1♁ same locality and collector, 16–31 OCT 2019 (ANIC); 1♁ same locality and collector, 1–15 MAY 2020 (ANIC); 1♁ same locality and collector, 16–31 OCT 2020 (ANIC); 3♁♁ same locality and collector, 16–31 DEC 2020 (ANIC); 2♁♁ same locality and collector, 16–31 JAN 2021 (ANIC); 1♁ same locality and collector, 16–28 FEB 2021 (ANIC); 1♁ same locality and collector, 16–30 NOV 2021 (ANIC); 1♁ same locality and collector, 1–15 DEC 2021 (ANIC); 1♁ same locality and collector, 16–31 DEC 2021 (ANIC); 1♁ same locality and collector, 1–15 FEB 2022 (ANIC); 2♁♁ same locality and collector, 1–15 MAR 2022 (ANIC); 1♁ Whitfield Ra., SW. of Cairns 520m, 14 DEC 1970, J.G. Brooks, at light (ANIC); 1♁ Isley Hills 1050m, 17°03’S 145°42’E, 30 NOV 1993, Cook, Monteith & Janetzki (QM); 1♀ nymph 1♁ North Bell Peak, Malbon Thompson Ra. 800–1000m, 19–22 NOV 1990, Monteith & Thompson (QM); 1♀ Nth. Bell Peak, 20 km S Cairns 900–1000m, 15–16 SEP 1981, G. Monteith & D. Cook (QM); 1♀ with ootheca, Lake Eacham 17.17°S 145.38°E, 8 FEB 1989, G. Milledge, MAN-44 (MV); 1♁ Lake Barrine, Atherton Tableland, 17.15 S 145.38 E, 11 DEC 1985, J. Balderson, Stop A-8 (ANIC); 1♀ Massey Range, 4km W of Centre Bellenden Ker 1250m, 17°16’S 145°49’E, 9–11 OCT 1991, Monteith, Janetzki & Cook (QM); 2♁♁ Mt Baldy Loop Rd, nr Atherton Herberton Range 1097m, 17.16’S 145.25’E. (GPS), 20 MAR 2010, D.C.F. Rentz & B. Richardson, Stop 3, ANIC Image Database No. 11 000024–000025 (ANIC); 1♁ Mt Baldy, 17.27 S 145.42 E, 23 MAR 2022, M.G. Connors, C. Henderson & M. Allan, at lights (ANIC); 1♀ same locality, date, and collectors, on tree trunk with lichen (ANIC); 1♀ nymph, same locality, date, and collectors, on tree trunk with lichen, raised in captivity, died 11 MAY 2022 (ANIC); 1♀ same locality, date, and collectors, on tree trunk with lichen, raised in captivity, died 19 NOV 2022 (ANIC); 1♀ Curtain Fig, 2km SSW. of Yungaburra, Atherton Tableland, 17.17 S 145.34 E, 13 DEC 1985, J. Balderson (ANIC); 1♁ Wongabel St For., forest track off Pavey Rd 746m, 17°19.112 S 145°30.315 E (car), 26 NOV 2019, D.C.F. Rentz, Stop 28 (ANIC); 2♀♀ nymphs, Wongabel, 17.332 S 145.503 E, 30 APR 2022, M.G. Connors, C. Henderson & M. Allan, on tree trunk with moss (ANIC); 1♀ with ootheca, same locality, date, and collectors, collected as nymph and raised in captivity, matured JUL 2022 (ANIC); 1♁ 9km W. of Crawfords Lookout (1 road km S. of Palmerston Hwy.), Palmerston Nat. Park, 17.35 S 145.43 E, 11 FEB 1988, D.C.F. Rentz, at light, Stop A-16 (ANIC); 1♀ Mt. Fisher 7km SW Millaa Millaa (Kjellberg Rd) 1000m, 3 MAY 1983, G.B. Monteith & D.K. Yeates (QM); 1♀ Palmerston Nat. Park, 12km SE. of Millaa Millaa, 17.35 S 145.42 E, 15 DEC 1985, J. Balderson (ANIC); 1♀ Palmerston Nat. Pk. 350–400m, 2 JAN 1990, G.B. Monteith (QM); 1♀ Hinchinbrook Is, Upper Gayundah Ck 850m, 10 NOV 1984, G. Monteith, pyrethrum knockdown (QM); 1♁ Paluma, 1 OCT 1979, D. Frith, light trap in rainforest, Genitalia prep. MG335 J. Balderson (ANIC); 1♁ same locality and collector, DEC 1980, at light (ANIC); 1♁ Mt Halifax summit, 21 MAR – 10 MAY 1991, D. Cook, heath, pitfalls & intercepts (QM); 1♁ Mt Elliott summit 1150m, 26 MAR – 12 MAY 1991, D. Cook, rainforest pitfall & intercepts (QM). Differential diagnosis. C. oxynota can be distinguished from C. australis and C. paraoxypila by the keeled dorsomedian lobes on the third to sixth abdominal tergites of the male abdomen reaching approximately one fifth the length of the following tergite, by the expanded but short, flattened dorsomedian lobes on the third to fifth tergites of the female abdomen, and by the L4A of the male genitalia possessing a dextrally-directed, rounded lobe on the right side. Additionally, C. oxynota can be distinguished from C. australis by the paired tubercles dorsolaterally on the anterior metazone, and it can be distinguished from C. paraoxypila by the slender body form, lack of a postocellar process, and short female tegmina that almost reach the hind margin of the first abdominal tergite. Description. Head. Head slightly longer in female. Clypeus with anterior area depressed; anterior margin slightly sinuate in male, more sinuate in female. Lower frons generally six-sided, five-sided in some females. Eyes distinctly projecting dorsally in female. Ocellar tubercle prominent, weakly raised in male, in female low and indistinct; in male ocelli moderately large, ovate, median ocellus distinctly smaller than lateral ocelli, in female all ocelli very small, almost absent. Vertex without obvious postocellar process but broadly raised behind ocelli, weakly concave in dorsal view; juxtaocular bulge weak in male, not strongly projecting beyond vertex, in female stronger, projecting beyond eyes and vertex (Figure 5D–E). Antennae reaching beyond midpoint of pronotum in female. Thorax. Pronotum elongate, somewhat slender, approximately 2.3–2.5 times as long as wide in male, approximately 2.0–2.3 times as long as wide in female, broadest just posterior to supracoxal sulcus. Median keel well-defined on almost entire metazone, generally slightly stronger in female. Prozone ovate, with sides almost parallel, anteriorly with a broad elevated tubercle in male, in female moderately elevated anteriorly; posteriorly with low, undulating diagonal ridges that extend onto lateral prozone. Metazone elongate, approximately 1.7 times length of prozone, anterodorsally with a pair of widely-spaced dorsolaterally-projecting tubercles, these stronger in female, dorsolateral longitudinal carinae rather long, covering posterior half except extreme posterior region, posterior tubercle with a pair of blunt, dorsally-directed spines or rounded knobs in male, in female with a pair of elongate, blunted, dorsally-directed spines. Posterolateral expansion usually acutely-angled and somewhat expanded adjacent to anterior corners of metazone (Figure 5A–B, 6A–B). Postcervical sclerite with anterior margin broadly incised; T-shaped sclerite somewhat narrow medially. Foreleg spination formula: F = 3DS/7–12AvS/4PvS; T = 4–8AvS/4–7PvS. A single aberrant but apparently undamaged individual from Lake Barrine has only 3 posteroventral forefemoral spines and a strongly reduced middle discoidal spine on the right forefemur. Legs. Forecoxa very elongate, narrow. Forefemur elongate, narrow, much more so in male, broadest slightly distal to tibial spur groove, concave dorsally; posterior keel very weak, sometimes almost absent; tibial spur groove weakly-defined distally; anteroventral forefemur spines alternating between small and large, with the distal four spines iiiI, Iii

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