Abyssorchomene abyssorum

Abyssorchomene abyssorum (Stebbing, 1888) Figs 2–9 Orchomene abyssorum Stebbing, 1888: 676–679, pl. 21. Orchomenopsis abyssorum Sars 1895: 74. — Stebbing 1906: 84, fig. 14. —? Chevreux 1900: 23–24. — Chevreux 1903: 92. —? Walker 1903: 224, 232. — Chevreux 1905: 7 (in list). —? Stephensen 1925: 125. — Chevreux 1935: 59–60 (in part, part = Abyssorchomene cf. abyssorum). Anonyx abyssorum Della Valle 1893: 824. Orchomenella abyssorum Ruffo 1949: 10 (in list). – Schellenberg 1955: 192 (in list). — Barnard J.L. 1958: 96 (in list). —? Birstein & Vinogradov 1960: 188–189, 227, 232, fig. 8. — Gurjanova 1962: 433 (in key). — Hurley 1963: 125–126. — Barnard J.L. 1964: 86, 89 (in key). —? Birstein & Vinogradov 1964: 164. — Thurston & Allen 1969: 364. — Sanderson 1973: 37 (in list). —? Lowry & Stoddart 1994: 129, 180–181. — Vinogradov G. 1999: 1147, 1164–1165 (in part; part = Abyssorchomene scotianensis). Orchomene abyssorum – Lowry & Bullock 1976: 94–95 (in part; part = Abyssorchomene scotianensis). — Shulenberger & Barnard 1976: 248. — Andres 1983: 209, 211–212. —? Austin 1985: 601 (in list).— Barnard & Karaman 1991: 508 (in part; part = Abyssorchomene scotianensis). —? Kaufmann 1992: 1–170. —? Bellan-Santini 1998: 145–146, 148. Abyssorchomene abyssorum – De Broyer 1983: 142–144 (in part; part = Abyssorchomene scotianensis). — Palerud & Vader 1991: 32 (in list). —? Jones et al. 1998: 1124–1125. —? Witte 1999: 143. —? Janssen et al. 2000: 3005, 3010–3011, 3013, 3015. — Thurston 2001: 358, 360, 362, 365, 369 (in part; part = Abyssorchomene scotianensis). —? Treude et al. 2002: 1284–1285, 1288. —? Horton 2005: 1. — Brandt et al. 2012: 144, 146, 152, 155 (biogeography). (in part). — Lowry & Kilgallen 2014: 6–9. —? Duffy et al. 2016a: 1691–1692, 1694–1696. —? Duffy et al. 2016b: 424. —? Lacey et al. 2016: 126, 128, 131. — Corbari & Sorbe 2018: 2. —? Bribiesca-Contreras et al. 2021: 9. Orchomenopsis (Orchomene) abyssorum – Costello et al. 1989: 32 (in list). Orchomene (Abyssorchomene) abyssorum –? Barnard & Ingram 1990: 26–29, figs 15–17. non Orchomenopsis chilensis f. abyssorum – Schellenberg 1926: 291–292, fig. 27 (= Abyssorchomene scotianensis). non Orchomenella abyssorum – Barnard K.H. 1932: 69, fig. 28 (= Abyssorchomene cf. scotianensis, except fig. 27b = Abyssorchomene abyssorum). — Nicholls 1938: 35, fig. 15 (= Abyssorchomene cf. scotianensis). — Dahl 1954: 282 (= Abyssorchomene cf. scotianensis). — Dahl 1959: 225 (= Abyssorchomene sp. nov.). — Birstein & Vinogradov 1962: 41 (= Abyssorchomene cf. scotianensis). non Orchomene abyssorum Arnaud 1974: 572 (ecology), (= Abyssorchomene scotianensis). — Lowry 1982: 320 (= Abyssorchomene scotianensis). — Wakabara et al. 1990: 2, 4, 6 (= Abyssorchomene cf. scotianensis). non Abyssorchomene abyssorum – Thurston 1990: 262–263, 269 (= A. patriciae in part; part = A. cf. patriciae; T. Horton, pers. com.). — Diffenthal & Horton 2007: 31 (= A. gerulicorbis; T. Horton, pers. com.). — Horton & Thurston 2009: 433–434 (= A. gerulicorbis; T. Horton, pers. com.). — Gutteridge 2012: 5, 22, 24 (= A. patriciae; T. Horton, pers. com.). — Corrigan et al. 2013: 156, 158–161 (= A. cf. patriciae). — Cousins et al. 2013: 303-304 (= A. cf. shannonae; T. Horton, pers. com.). — Duffy et al. 2013: 360–368 (= A. patriciae; T. Horton, pers. com.). — Horton et al. 2013: 352, 354–358 (= A. patriciae; T. Horton, pers. com.). — Priede et al. 2013: 8 (= A. cf. patriciae). — Horton et al. 2020: 6–7, 11 (= A. patriciae; T. Horton, pers. com.). New diagnosis Pereonites 1–7 and pleonites 1–2 with a weak but distinct dorsoposterior hump on each segment. Lateral cephalic lobe broadly rounded, dorsal margin regularly convex and slightly more convex than the nearly straight ventral margin. Antennae 1–2 of male with calceoli, female without. Epistome level with upper lip. Maxilla 1 palp, distal end weakly convex with conical apical spines contiguous. Maxilliped inner plate, distal margin conspicuously excavate; outer plate inner margin not scalloped. Coxa 1 slightly widened distally, ~ 1.2–1.3× proximal width. Gnathopod 2 propodus of male and female narrow, length ~ 2.3–2.5 × width, dactylus very small, inserted on the posterodistal one-third of distal margin, palm lacking concavity. Coxa 5 very weakly posterolobate, posterodistal lobe narrow with posterodistal margin nearly straight. Pereopod 7 basis with anterior margin slightly concave, posterior margin subparallel to anterior margin in the proximal two-thirds, distal third with a distinct, nearly straight bevel (male less distinct and more rounded). Uropod 1 peduncle length greater than 1.5–1.7 × length of outer ramus. Uropod 3 inner ramus extends past distal end of article 1 and reaches ~ 70% length of article 2 of outer ramus. Telson cleft ~ 52–56% of length. Material examined Holotype SOUTHWEST ATLANTIC • ♂ (7.5 mm, figured, see Figs 5–9, dissected in 3 tubes, carcass missing/lost); east of Buenos Aires; HMS Challenger, station 323; 35°39′ S, 50°47′ W; depth 3475 m (1900 fathoms); bottom blue mud, temperature 33.1°F; 28 Feb. 1876; BMNH 1889.5.15.23 Additional material SOUTHWEST ATLANTIC • 1 ♀ (9.5 mm, mature, with setose brood plates, illustrated, see Figs 2–4, appendages on 2 slides); Argentine Basin; RV Meteor DIVA-3 M79/1 expedition, station 531; 35°56.49′ S, 48°53.85′ W; depth 4586 m; gear, baited traps; 15 Jul. 2009; E. Hendrycks leg.; ZMH K-61206 • 1 ♂ (7.0 mm, appendages on 2 slides, partly illustrated, see Fig. 2); same collection data as for preceding; ZMH K-61207 • 5 ♂♂, 5 ♀♀, 4 juveniles; same collection data as for preceding; ZMH K-61208 • 10 ♂♂, (one of these 7.0 mm, dissected, appendages on 3 slides), 5 ♀♀, 7 juveniles; same collection data as for preceding; CMNC 2022-0001 • 5 ♂♂, 10 ♀♀, 3 juveniles; same collection data as for preceding; RBINS INV. 138.486. NORTH ATLANTIC • 9 ♀♀ (with mature ♀♀, 7–10 mm, 1 mature ♀, 10 mm, appendages on 8 slides, 1 nearly mature ♀, 9.5 mm, appendages on 3 slides); near Azores; Hirondelle 1896 campaign, station 730; 37°58′ N, 28°33′30″W; depth 2660 m; bottom sandy clay; gear, baited traps; 3–5 Aug. 1896; MNHN, collection Chevreux (no registration number, see Chevreux 1903: 92). Description Female Based on female (mature), 9.5 mm (ZMH K-61206). PEREONITES 1–7 AND PLEONITES 1–2 (Fig. 2). With weak but distinct dorsoposterior hump on each segment. PLEONITE 3 (Fig. 2). With a distinct, rounded posterodorsal elevation slightly overhanging urosomite 1. COXAE 1–2 (Figs 2–3). Subequal to slightly longer (1.1 × to 1.25 ×) than the depth of corresponding pereonites (in lateral view). COXAE 3–4 (Fig. 4). Slightly longer (1.1 × to 1.25 ×) than the depth of corresponding pereonites, coxa 3 not shown. EPIMERON 3 (Fig. 2). Subquadrate, with posterodistal angle slightly obtuse, apex rounded, posterior margin weakly but regularly convex, ventral margin regularly convex. UROSOMITE 1 (Fig. 2). With a deep, narrow dorsal concavity in front of the strong, regularly rounded dorsal boss, convex on posterior margin, boss projecting strongly upright and backward, slightly overhanging urosomite 2. HEAD (Fig. 2). About equal in length to pereonite 1. LATERAL CEPHALIC LOBE (Fig. 2). Broadly rounded; dorsal margin regularly convex and slightly more convex than the nearly straight ventral margin. EYE (Fig. 2). Nearly indistinguishable, but non ommatidial, formed of pigment granules; crescent-shaped, narrow, parallel to head anterior margin (estimated in figure by dotted outline). ANTENNA 1 (Fig. 2). Peduncular article 1 without anterodistal lobe, slightly dilated, length about 1.4× width; flagellum 10-articulate, first article of flagellum callynophorate, densely furnished with double row of aesthetascs medially, callynophore small, shorter (67%) than the remaining articles combined; accessory flagellum 5-articulate, first article slightly shorter (80%) than remaining articles combined; calceoli absent. ANTENNA 2 (Fig. 2). Slightly longer than antenna 1; geniculate between peduncular articles 3–4, peduncular articles 4–5 lined with anteromedial brush setae; flagellum 12-articulate, calceoli absent. EPISTOME (Fig 2). Level with the upper lip, from which it is separated by a small slit and weak shallow indentation; weakly concave above upper lip. UPPER LIP (Fig. 2). Anterior margin convex, not protruding; midventral margin with a small, angular projection. MANDIBLE. Incisor distinctly convex and slightly widened, dorsolateral and ventromedial corners with small tooth; left lacinia mobilis slender, curved, distally with small teeth, right lacking; accessory spine row with 3 strong spines, interspersed with fine setae; molar somewhat falciform, not columnar, forming a narrow crest, acutely produced on proximal end, setose with small triturative surface, hairy process (cf. Oleröd 1975) located proximal to molar; palp attached proximal to molar; article 2 with 15A2 setae, article 3 missing (refer to Fig. 6 of male holotype for general morphology and setae details). LOWER LIP. Outer lobes broad, inner margins strongly setose, distal inner margins excavated; without inner lobes; mandibular processes rounded. MAXILLA 1 (Fig. 2). Outer plate with 11 spine-teeth in 7/4 crown arrangement; palp article 2 widened, with 9 contiguous (or nearly contiguous, separated by much less than the width of a spine) distal conical spines and one apical strong seta on outer corner; inner plate short with subrounded apical projection, and two unequally sized plumose setae inserted subapically (refer to Fig. 7 of male holotype for general morphology). MAXILLA 2. Outer and inner plates strongly tapering distally, both lined with strong rows of pectinate medial marginal spines and setae; inner plate much shorter and slightly narrower than outer plate, apex just reaching the proximal end of medial setal row of outer plate (refer to Fig. 7 of male holotype for general morphology). MAXILLIPED (Fig. 2). Inner plate short, reaching about one third length of outer plate and reachi

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