Parapterogramma gonitellum Kuwahara, Marshall & Luk, 2025, sp. nov.

Parapterogramma gonitellum sp. nov. urn:lsid:zoobank.org:act: 774B9202-A4C3-4816-BD47-3266E6C0F427 Figs 11– 13, 40E, 42 Etymology This name refers to the highly reduced male postgonites. Material examined Holotype AUSTRALIA – Queensland • ♂; Atherton Tablelands, Lake Eacham, Chambers Lodge; 6–8 Apr. 1999; S.A. Marshall leg.; dung; QMBA debu00163357. Paratypes AUSTRALIA – Queensland • 3 ♂♂, 10 ♀♀; same collection data as for holotype; DEBU • 2 ♂♂, 13 ♀♀; same collection data as for holotype; QMBA • 10 ♂♂, 8 ♀♀; Fitzroy Island, site 2; 16°56′ S, 145°59′ E; 40 m a.s.l.; 19–21 Apr. 1999; G.B. Monteith leg.; transition to rainforest, dung trap; DEBU • 12 ♂♂, 10 ♀♀; same collection data as for preceding; QMBA • 4 ♂♂, 15 ♀♀; same collection data as for preceding; site 4; 15 m a.s.l.; QMBA • 3 ♂♂, 4 ♀♀; Lake Euramoo; 17°10′ S, 145°36′ E; 700 m a.s.l.; 10–11 Feb. 1999; D.J. Cook leg.; rainforest, dung pitfall; DEBU • 4 ♂♂, 4 ♀♀; same collection data as for preceding; QMBA • 7 ♂♂, 3 ♀♀; 3.5 km SE of Malanda; 17°23′ S, 145°36′ E; 760 m a.s.l.; 6–9 Feb. 1999; Monteith and Cook leg.; rainforest, dung pitfall; DEBU • 8 ♂♂, 4 ♀♀; same collection data as for preceding; QMBA. Other material examined AUSTRALIA – Queensland: • 1 ♂, 1 ♀; Above Mill Creek; 17°30′ S, 145°27′ E; 1030 m a.s.l.; 5–9 Feb. 1999; Monteith and Cook leg.; open forest, fungus pitfall; QMBA • 3 ♂♂, 3 ♀♀; Atherton Tablelands, Lake Eacham; 5–8 Apr. 1999; S.A. Marshall leg.; sweep; DEBU • 1 ♀; Baldy Mountain Road, start; 17°21′ S, 145°25′ E; 860 m a.s.l.; 3–6 Feb. 1999; Monteith and Cook leg.; open forest, fish pitfall; QMBA • 1 ♀; same collection data as for preceding; fungus pitfall; QMBA • 1 ♀; 1 km NW of Buchan Point; 16°44′ S, 145°39′ E; 10 m a.s.l.; 3–11 Feb. 1999; Monteith and Cook leg.; open forest, dung pitfall; QMBA • 2 ♂♂; same collection data as for preceding; fish pitfall; QMBA • 4 ♂♂, 2 ♀♀; Buchan Point; 16°44′ S, 145°39′ E; 5 m a.s.l.; 11–13 Feb. 1999; G.B. Monteith leg.; rainforest, dung pitfall; QMBA • 3 ♀♀; Cape Tribulation; 10 m a.s.l.; 15 Jul. 1982; S. Peck leg.; rainforest litter, carrion trap; DEBU • 2 ♀♀; Cape Tribulation, 40 km N of Daintree; 10 m a.s.l.; 12–17 Jul. 1982; S. Peck leg.; rainforest, carrion trap; DEBU • 3 ♀♀; Cheelonga Creek, Evelyn; 17°31′ S, 145°32′ E; 1080 m a.s.l.; 15–17 Apr. 1999; G.B. and S.R. Monteith leg.; rainforest, dung trap; QMBA • 4 ♀♀; Cooktown, Keatings Gap; 100 m a.s.l.; 28–31 Jun. 1982; S. and J. Peck leg.; rainforest, dung trap; DEBU • 2 ♂♂; 4 km S of Hartley’ s Creek 16°41′ S, 145°35′ E; 5 m a.s.l.; 3–11 Feb. 1999; Monteith and Cook leg.; littoral rainforest, dung pitfall; QMBA • 2 ♂♂, 1 ♀; same collection data as for preceding; fish pitfall; QMBA • 8 ♂♂, 6 ♀♀; same collection data as for preceding; fungus pitfall; QMBA • 2 ♂♂; 1 km SW of Kauri Creek; 17°09′ S, 145°35′ E; 700 m a.s.l.; 10–11 Feb. 1999; D.J. Cook leg.; rainforest, dung pitfall; QMBA • 8 ♂♂, 5 ♀♀; 2 km E of Kauri Creek; 17°08′ S, 145°37′ E; 680 m a.s.l.; 10–11 Feb. 1999; Monteith and Cook leg.; rainforest, fungus pitfall; QMBA • 2 ♀♀; Kjellberg Road, State Forest boundary; 17°32′ S, 145°35′ E; 920 m a.s.l.; 6–9 Feb. 1999; Monteith and Cook leg.; rainforest, dung pitfall; QMBA • 6 ♂♂, 5 ♀♀; same collection data as for preceding; fungus pitfall; QMBA • 7 ♂♂; Kjellberg Road turnoff; 17°32′ S, 145°36′ E; 740 m a.s.l.; 6–9 Feb. 1999; Monteith and Cook leg.; rainforest, fungus pitfall; QMBA • 4 ♂♂, 13 ♀♀; Lacey’ s Creek, 10 km SE of El Arish, near Mission Beach; 40 m a.s.l.; 23 Jun.–5 Aug. 1982; S. and J. Peck leg.; rainforest, flight intercept trap; DEBU • 5 ♂♂; Maalan State Forest, Ravenshoe Road; 17°35′ S, 145°36′ E; 840 m a.s.l.; 6–9 Feb. 1999; Monteith and Cook leg.; rainforest, fungus pitfall; QMBA • 1 ♂; Mill Creek; 17°30′ S, 145°27′ E; 940 m a.s.l.; 5–9 Feb. 1999; Monteith and Cook leg.; open forest, fungus pitfall; QMBA • 1 ♀; Mill Creek, above; 17°30′ S, 145°27′ E; 1000 m a.s.l.; 5–9 Feb. 1999; Monteith and Cook leg.; open forest, rotten bait; QMBA • 1 ♂, 1 ♀; Millaa-Millaa Falls; 17°30′ S, 145°37′ E; 800 m a.s.l.; 6–9 Feb. 1999; Monteith and Cook leg.; rainforest, fungus pitfall; QMBA • 4 ♂♂, 7 ♀♀; Millstream Conservation Park; 17°32′ S, 145°29′ E; 1040 m a.s.l.; 3–5 Feb. 1999; Monteith and Cook leg.; wet sclerophyll, dung trap; QMBA • 3 ♂♂, 7 ♀♀; same collection data as for preceding; fish pitfall; QMBA • 19 ♂♂, 10 ♀♀; same collection data as for preceding; fungus trap; QMBA • 1 ♀; Millstream Conservation Park, adjacent; 17°32′ S, 145°29′ E; 1040 m a.s.l.; 4–5 Feb. 1999; G.B. Monteith leg.; open forest, automatic dung trap (day); QMBA • 1 ♂; Millstream Retreat; 17°32′ S, 145°31′ E; 1020 m a.s.l.; 14–17 Apr. 1999; G.B. and S.R. Monteith leg.; wet sclerophyll, fungus trap; QMBA • 3 ♂♂, 3 ♀♀; The Millstream, 10 km NNE of Ravenshoe; 17°32′ S, 145°31′ E; 1040 m a.s.l.; 3–5 Feb. 1999; Monteith and Cook leg.; rainforest, fish pitfall; QMBA • 1 ♂, 1 ♀; same collection data as for preceding; dung trap; QMBA • 2 ♂♂; The Millstream, 10 km NNE of Ravenshoe; 17°32′ S, 145°31′ E; 1040 m a.s.l.; 6 Dec. 1998 – 5 Feb. 1999; Cook and Monteith leg.; rainforest, pitfall trap; QMBA • 2 ♂♂, 5 ♀♀; The Millstream, 10 km NNE of Ravenshoe; 17°32′ S, 145°31′ E; 1050 m a.s.l.; 14–17 Apr. 1999; G.B. and S.R. Monteith leg.; wet sclerophyll, dung trap; QMBA • 6 ♂♂, 5 ♀♀; same collection data as for preceding; fungus trap; QMBA • 1 ♂, 1 ♀; Mount Finnigan Slopes, 30 km S of Cooktown; 400 m a.s.l.; 2–5 Jul. 1982; S. and J. Peck leg.; rainforest, carrion trap; DEBU • 1 ♀; same collection data as for preceding; litter and fungi; 1 Jul. 1982; DEBU • 1 ♂; Mount Hugh Nelson, south base; 17°31′ S, 145°33′ E; 1080 m a.s.l.; 7–9 Feb. 1999; G.B. Monteith leg.; rainforest, dung pitfall; QMBA • 4 ♂♂, 1 ♀; same collection data as for preceding; fungus pitfall; QMBA • 1 ♂, 1 ♀; same collection data as for preceding; rotten bait; QMBA • 22 ♂♂, 10 ♀♀; Mount Hypipamee National Park; 11–17 Apr. 1999; S.A. Marshall leg.; dung; DEBU • 3 ♂♂, 2 ♀♀; Oak Beach; 16°36′ S, 145°31′ E; 5 m a.s.l.; 3–11 Feb. 1999; Monteith and Cook leg.; rainforest, fish pitfall; QMBA • 7 ♂♂, 3 ♀♀; same collection data as for preceding; fungus pitfall; QMBA • 1 ♂, 2 ♀♀; Palmerston National Park, near Innisfail; 360 m a.s.l.; 2–5 Aug. 1982; S. and J. Peck leg.; carrion trap; DEBU • 1 ♀; 3 km SW of Ravenshoe; 17°38′ S, 145°28′ E; 860 m a.s.l.; 3–5 Feb. 1999; Monteith and Cook leg.; open forest, fungus pitfall; QMBA • 2 ♂♂, 1 ♀; Ravenshoe, 4 km SSE; 17°39′ S, 145°30′ E; 930 m a.s.l.; 4–6 Feb. 1999; Monteith and leg.; open forest, fungus pitfall; QMBA • 1 ♀; 7 km NE of Ravenshoe; 17°34′ S, 145°31′ E; 1080 m a.s.l.; 6 Dec. 1998 – 5 Feb. 1999; Monteith and Cook leg.; wet sclerophyll, pitfall; QMBA • 1 ♀; 9 km N of Ravenshoe; 17°32′ S, 145°29′ E; 1060 m a.s.l.; 3–5 Feb. 1999; Monteith and Cook leg.; wet sclerophyll, dung pitfall; QMBA • 1 ♀; same collection data as for preceding; fish pitfall; QMBA • 1 ♂, 1 ♀; same collection data as for preceding; fungus pitfall; QMBA • 1 ♂; 9.5 km N of Ravenshoe; 17°32′ S, 145°29′ E; 1060 m a.s.l.; 14–17 Apr. 1999; G.B. and S.R. Monteith leg.; wet sclerophyll, dung trap; QMBA • 1 ♀; Rosina Creek, 14 km SE of Millaa Millaa, Palmerston Highway; 720 m a.s.l.; 24 Jun.–2 Aug. 1982; S. Peck leg.; rainforest, flight intercept trap; DEBU • 1 ♂, 1 ♀; Shipton’ s Flat; 15°48′ S, 145°14′ E; 220 m a.s.l.; 19–22 Nov. 1998; G.B. Monteith leg.; rainforest, automatic dung trap (day); QMBA • 1 ♂, 4 ♀♀; Sluice Creek; 17°32′ S, 145°32′ E; 1100 m a.s.l.; 14–17 Apr. 1999; G.B. and S.R. Monteith leg.; rainforest, fungus trap; QMBA • 3 ♀♀; Thorton Range, near Daintree; 200 m a.s.l.; 12–18 Jul. 1982; S. and J. Peck leg.; carrion trap; DEBU • 9 ♂♂, 1 ♀; Tolga Scrub; 17°15′ S, 145°29′ E; 760 m a.s.l.; 3–6 Feb. 1999; Monteith and Cook leg.; rainforest, fungus pitfall; QMBA • 1 ♂; Vine Creek, Majors Mountain; 17°41′ S, 145°32′ E; 1060 m a.s.l.; 4–6 Feb. 1999; Monteith and Cook leg.; rainforest, fungus pitfall; QMBA • 1 ♂; 2 km SSE of Wondecla; 17°26′ S, 145°24′ E; 910 m a.s.l.; 3–6 Feb. 1999; Monteith and Cook leg.; open forest, dung pitfall; QMBA • 8 ♂♂, 13 ♀♀; Wonga Beach, 10 km S of Daintree; 15–16 Apr. 1999; S.A. Marshall leg.; low forest, dung; DEBU. Description BODY (Fig. 11). Length 1.2–1.5 mm. Head caramel-brown, darkened around lateral and posterior margins, with pale yellow frontal vitta and interfrontal plates, and silvery-gold microtomentum on ocellar triangle, orbital plates and occiput; gena yellow; antenna brown. Frontal width 2.0 × interfrontal height. Eye height 2.5× genal height. Thorax brown, anepimeron with a pale ventral spot. Scutum with pale grey and brown microtomentum forming two dark longitudinal stripes, lateral margins broadly yellow microtomentose, pale M-shaped microtomentose mark at level of posterior dorsocentral bristles; scutellum dark brown with median, grey-brown microtomentose triangle. Anterior dorsocentral bristles separated by 7–8 rows of acrostichal setulae. Legs brown, tibiae proximally paler, tarsi paler. Male mid tibia with a stout apicoventral bristle and two rows of stout ventral setae. Wing (Fig. 40E) pattern strong, anterior edge with a single darker spot below R 1. CS2 0.4 × CS3. R 2+3 gently sinuate, distal curve much stronger than basal curve, angled at ~60° to costa and broadly separated from costa for most of its length; costa extending well beyond apex of R 4+5 (4–5 × costal width). M 1 sometimes extending slightly beyond dm-m; M 4 indistinct beyond dm-m; CuA+CuP short and non-tubular. Halter off-white. MALE ABDOMEN (Fig. 12). S5 narrow, posteromedially desclerotized with long posterolateral setae. S6 sinuate. Cercus flat, triangular with several enlarged setulae, and widely separated between the pair; subepandrial sclerite semicircular, anteromedially darkened and posteromedially emarginate. Surstylus bilobed, the lobes almost entirely separated: anterior lobe somewhat C-shaped, with a broad basal half and a pointed apical half that curves underneath the b

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