Iteaphila longipalpis Sinclair & Shamshev 2021, comb. nov.

Iteaphila longipalpis (Melander) comb. nov. (Figs 8, 48, 64) Anthepiscopus longipalpis Melander, 1928: 106. Type locality: Arlington, Washington, USA. Anthepiscopus longipalpis: Melander, 1965: 455. Type material examined. HOLOTYPE ♀, labelled: “Arlington [48°10′52″N 122°8′20″W]/ 5.21.[190]8Was”; “window”; “ TYPE / Anthepiscopus / longipalpis/ Mel. [red label]”; “ALMelander/ Collection/ 1961” (USNM). Additional material examined. CANADA. Alberta: Banff NP [51.5°N 116.0°W], 14 mi W Banff, Ledum groenlandicum, 7.vii.1955, GES (1 ♀, CNC); Banff NP, 26 km N Lk. Louise, trib. Mosquito Ck [51°37′N 116°19′W], 28.vi.1989, BJS (1 ♂, CNC). British Columbia: 37 km W Anahim Lake [52°24′N 125°14′W], 6.vii.1992, A. Borkent (2 ♂, CNC); same data except, 48 km NW, 14.vii.1992 (1 ♂, CNC); Cowichan Lk. [48°49′N 124°02′W], 19.vi.1964, J.A. Clapman (2 ♀, CNC); Cultus Lk. [49°04′N 121°58′W], 24.v.1965, F. Schmid (1 ♂, 1 ♀, CNC); Galiano Is. [48°55′N 123°25′W], Spanish Hills, 29.iv.1979, 9.v.1982, G.G.E. Scudder (3 ♀, UBCZ); Kilimat [54°03′17″N 128°39′28″W], 2.vi.1960, JGC (2 ♀, CNC); same data except, 30.vi.1960, GES (1 ♂, CNC); King Salmon Lk., 58°43′N 132°54′W, 15.vii.1960, R. Pilfrey (1 ♀, CNC); Kitsumkalum Lk. [54°43′22″N 128°46′34″W], 20 mi N Terrace, 31.v., 16.vi.1960, JGC (1 ♂, 1 ♀, CNC); Kleanza Ck [54°36′05″N 128°24′11″W], 14 mi E Terrace, 17.vi., 4.vii.1960, CHM (2 ♀, CNC); same data except, 19.vii.1960, W. R. Richards (2 ♀, CNC); same data except, 29.vi., 4.vii.1960, JGC (1 ♀, CNC); same data except, 17.vi.1960, B. Pilfrey (1 ♀, CNC); Mission City [49°2′30″N 122°14′41″W], 3 and 9.vi.1953, W. R.M. Mason (2 ♂, 1 ♀, CNC); Mt. Allard, 23 mi N Terrace, 11.viii.1960, CHM 1 ♀, CNC); Mt. Thornhill, nr Terrace [54°30′44″N 128°32′18″W], 21.vi., 14.vii.1960, JGC (1 ♂, 7 ♀, CNC); same data except, 14 and 26.vii.1960, B. Heming (1 ♂, 1 ♀, CNC); same data except, 14.vii.1960, CHM (1 ♀, CNC); 8 km E Sicamous [50°53′N 118°54′W], 22.v.1992, AB (1 ♀, CNC); 24 km NE Sicamous, 19.v.1991, AB (1 ♀, CNC); 13 km SE Port Renfrew [48°30′N 124°18′W], 5.vii.1991, AB (4 ♀, CNC); Squamish, Diamond Head Tr. [49°45′N 123°03′W], 4000 ft, 28.viii.1953, GJS (1 ♀, CNC); same data except, 3500 ft, 5.viii.1953, E. Mason (1 ♂, CNC); 10 mi N Terrace, Glacier Ck, 7.vii.1960, JGC (7 ♀, CNC); 32 mi SW Terrace, 4–6.vi.1960, GES (2 ♀, CNC); same data except, 6.vi.1960, R. Pilfrey (1 ♀, CNC); Vancouver Is., Lk. Cowichan, 48°49′N 124°03′W, 28.vi.1983, J.A. Downes (1 ♀, CNC). Yukon: North Fork Crossing, mi 42 Peel Plateau Rd. [64°27′N 138°17′W], 3500 ft, 24.vi.1962, PJS (1 ♀, CNC). USA. Alaska: Isabel Pass [63°12′42″N 145°29′17″W], mi 206 Richardson Hwy, 2900 ft, 15.vi.1962, PJS (1 ♀, CNC); Thane [58°15′51″N 134°19′49″W], 1.viii.1952, WCF (1 ♀, WSU); White Pass [59°28′47″N 135°15′22″W], 10 mi Skagway, 30.vii.1996, PHA & MA (3 ♀, USNM). Oregon: Benton Co., Mary’s Peak Cpgd, Meadow Edge Tr., along Upper Parker Ck, 44°30′32.8″N 123°33′36.0″W, 1077 m, 4.vii.2014, JMC (1 ♂, CNC); Clackamas Co., Mt. Hood NF, Tollgate Cpgd [45°19′21″N 121°54′24″W], 22.vi.1977, DDW (8 ♀, CAS); Clackamas Co., Bruin Run Ck, Camp Ck Cpgd [45°18′16″N 121°52′02″W], 24.vi.1974, PHA (2 ♀, CAS); Clackamas Co., Mt. Hood NF, Still Ck Cpgd, 45°17.703′N 121°44.313′W, 1150 m, 9.vii.2013, BJS (1 ♀, CNC); Marion Co., Silver Falls SP, 23 and 24.vi.1974, PHA (7 ♀, CAS); Marion Co., Willamette NF, French Ck Rd., 44°45.969′N 122°11.211′W, 660 m, 10.vii.2013, BJS (1 ♀, CNC). Washington: Clallam Co., Olympic NF, Klahowya SP [48°03′52″N 124°06′41″W], Soleduck R., 30.vi.1974, PHA (1 ♀, CAS); Clallam Co., Olympic NP, meadow below Sunrise Ridge, Mt. Angeles [47°59′40″N 123°27′59″W], 1550 m, 22 and 24.vii.1978, DDW (4 ♂, CAS); same data except, Lake Creek, 24.vii.1978 (9 ♀, CAS); Clallam Co., Beaver Lk., 5 km NE Sappho, 48°06′51″N 124°14′40″W, 7.vi.2004, C.J. Borkent (4 ♀, CNC); Clallam Co., Olympic NP, nr. Fairholm Cpgd, 48°04′12″N 123°55′7.5″W, 24.vi.2005, C.J. Borkent (1 ♀, LEM); same data except, CNC161443, CNC161444, CNC161445, CNC161446 (1 ♂, 3 ♀, barcoded, LEM); Everett [47°58′45″N 122°12′06″W], 20.vi.1920, A Spuler (6 ♀, WSU); Everett, 19.vi.1920, 4.vii.1924, ALM (1 ♂, 27 ♀, USNM); Ilwaco [46°18′45″N 124°1′47″W], 5.v.1918, ALM (1 ♀, USNM); King Co., Seattle, Issaquah [47°32′8″N 122°2′36″W], 23.vii.1988, R. Danielsson (1 ♀, MZLU); King Co., 5.5 km W Greenwater, nr Federation Forest SP, Jct Hwy 410 & W-6300, 47°09′28″N 121°43′40″W, 515 m, 25.vi.2005, C.J. Borkent (1 ♀, LEM); Lewis Co., trib. Catt Ck, 2847 ft, 46°41′53″N 121°57′46″W, 16.vi.2018, JMC (1 ♂, 1 ♀, CNC); Lewis Co., stream at mi 9 Skate Ck Rd, 2460 ft, 46°43′32.01″N 121°49′51″W, 15.vi.2018, JMC (1 ♂, 2 ♀, CNC); Lewis Co., Knuppenburg Lk., 17 km ENE Packwood, Hwy 12, 1260 m, 46°37′46″N 121°24′57″W, 11.vi.2004, C.J. Borkent (1 ♀, LEM); Mason Co., 16 kmW Lilliwaup, Staircase, Olympic NP, N Fk Skokomish R., 47°31′06″N 123°19′58″W, 280 m, 10.vi.2004, C.J. Borkent (1 ♀, LEM); Mt. Constitution [48°40′39″N 122°49′52″W], 17.v.1910 (2 ♀, UNSM); Mt. Baker, Skyline Tr [48°52′51″N 121°51′52″W], 10.viii.1925, ALM (3 ♀, USNM); Mt. Rainier, Van Trump Ck [46°47′34″N 121°46′33″W],. ix.1917, ALM (1 ♀, USNM); Mt. Rainier NP, nr Cougar Cpgd [46°46′03″N 121°47′39″W], 8.vii.1989, BJS (2 ♀, CNC); Mt. Rainer NP, ck nr White River Cpgd [46°54′09″N 121°38′28″W], 1340 m, 3.viii.1977, DDW (2 ♂, CAS); same data except, Fish Ck, West Side Rd, 910 m, 1.viii.1977 (4 ♀, CAS); same data except, along Sunbeam Ck above Louise Lk. [46°46′16″N 121°43′23″W], 1520 m, 4.viii.1977 (1 ♀, CAS); Mt. Rainier NP, Narada Falls to Reflection Lk. Tr., 46°46′N 121°44′W, 1400 m, 17.vii.2012, BJS (1 ♂, 10 ♀, CNC); same data except, CNC197570, CNC197571, CNC197572 (2 ♂, 1 ♀, barcoded, CNC); same data except, ex. Erythronium montanum (1 ♂, CNC); same data except, Westside Rd nr. Fish Ck, 46°46.743′N 121°53.069′W, 880 m, 4.vii.2013 (1 ♂, 16 ♀, CNC); same data except, Stevens Ck Tr., 46°45.338′N 121°38.530′W, 1400 m, 17.vii.2012 (1 ♀, CNC); Mt. Rainier NP, Longmire Tr., marsh W Inn, 46°44′56″N 121°48′51″W, 850 m, 25.vi.2005, C.J. Borkent (2 ♀, LEM); Olympia [47°2′16″N 122°54′3″W], 22.vi.1920, ALM (1 ♂, 2 ♀, USNM); Olympic NP, Sol Due River [47°58′00″N 123°51′28″W], 6.vii.1978, D.E. Hardy (1 ♀, WSU); Pluvius [46°32′53″N 123°24′32″W], 10.vii.1917, ALM (1 ♀, USNM); Tacoma [47°14′29″N 122°27′34″W], 27.v.1917, ALM (1 ♂, USNM); Vashon [47°24′58″N 122°28′6″W], 28.v.1917, ALM (2 ♂, 11 ♀, USNM). Wyoming: Sheridan Co., Black Mtn [44°44′17″N 107°23′16″W], pine forest, ypans, dung, 5–20.vii.1990, J. Swann (1 ♂, 11 ♀, DEBU); Teton Co., Togowte Pass [43°45′00″N 110°04′48″W], 9000 ft, 22.vii.1953, R. R. Driesbach (1 ♂, CNC). Possible additional material. USA. California: Placer Co., E end Bear Valley [39°18′34″N 120°39′54″W], 1.vi.1964, PHA (2 ♀, CAS). Recognition. This species is distinguished by 4-serial acrostichal setae, thinly pruinescent scutum with dark chaetotaxy, elongate mouthparts, pale legs and features of the male terminalia (hypoproct process extending beyond cercus; narrow, upright surstylus) (Fig. 48). Redescription. Wing length 3.2–3.8 mm. Male (Fig. 8). Head dark brown in ground-colour, with pale brown to black setation, occiput thinly to densely greyish pollinose. Eyes holoptic, with upper ommatidia enlarged. Frons represented by small triangular space just above antennae, bare, greyish pollinose. Ocellar triangle prominent, with 2 pairs of long hair-like setae. Postvertical setae subequal to postocular setae, thin; occiput covered with numerous similar setae in lower part, pale brown posterior to mouthparts. Antenna dark brown; scape short, slightly shorter than globular pedicel, both with short setae; postpedicel rather narrow, nearly 4 times longer than basal width, base somewhat expanded and smoothly tapered; stylus ca 3–4 times longer than wide, sensillum-tipped; segment 9 tubular, 2–3 times longer than wide, apical sensillum about as long as segment 9. Proboscis very long, projected obliquely; labium longer than head height; palpus slender, projected parallel to labrum, somewhat shorter than labrum. Thorax dark brown in ground-colour with posterior part of postpronotal lobe and postalar tubercle often paler, with brown to black setation; scutum viewed dorsally entirely velvety brown, slightly matt, without vittae; viewed anteriorly densely pruinose; mesopleuron uniformly greyish brown pollinose. Proepisternum with a few hair-like setae. Postpronotal lobe with 1 long thin and several shorter setae. Mesonotal setae distinct, thin; acr 4-serial, arranged in 2 pairs of closely spaced irregular rows, lacking on prescutellar depression, distance between paired rows wider than length of acr; dc uniserial (with some additional setulae in anterior part), offset from row anteriorly, slightly longer than acr, 2–3 distinct prescutellar dc; 1 ph, 1 presut spal (sometimes with additional setulae), 3–4 npl (with a few additional shorter setae), several psut spal setulae, 1 pal and several short setulae, 6–7 pairs of sctl. Legs pale yellow to yellowish brown, coxae and tarsi somewhat darker. Coxae and trochanters with unmodified hair-like setae. Fore femur with row of short anteroventral hair-like setae, especially distinct apically; and somewhat longer setae on posteroventral, dorsal and posterior faces. Mid femur with moderately long posteroventral bristly setae along entire length, bearing some moderately long setae on dorsal and posterior faces. Hind femur with long anteroventral and dorsal setae along whole length. Tibiae with row of slightly prominent thin posterodorsal setae. Tarsomere 1 of mid and hindlegs with pair of rows of stiff ventral setae; tarsomere 5 not flattened on all legs; pulvilli broad, shorter than tarsal claw. Wing distinctly uniformly pale infuscate; basal costal seta absent; pterostigma distinct brownish, elliptical, overlapping apex of R 1; anal lobe very prominent, acute. Sc complete; R 2+3 often slightly bowed posterior to pterostigma; R 4+5 unbranched; cell dm broad, longer than bas

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