Anthomyza variegata

Anthomyza variegata (Loew, 1863) (Figs 381, 385–400, 437) Anthophilina variegata Loew, 1863: 324; OSTEN SACKEN (1878): 198 (catalogue). Anthomyza variegata: CZERNY (1902): 251 (key), 254 (list); ALDRICH (1905): 645 (catalogue); MELANDER (1913): 293 (key, distribution); SABROSKY (1965): 819 (catalogue); COLE (1969): 436 (distribution); ROHÁĆEK (1998a): 173 (checklist). Type material. LECTOTYPE: ♂ (designated herewith): “variegata m.” (Loew’s handwriting), “Loew Coll.”, “ Type 13428” (red label), “ LECTOTYPUS ♂ Anthophilina variegata Loew, J. Roháček & K. N. Barber des. 2013” (red) and “ Anthomyza variegata (Loew) ♂, J. Roháček & K. N. Barber det. 2013”. The specimen is in good condition, with clearly visible genitalia (see Fig. 381) (MCZC, intact). PARALECTOTYPE: ♀, “36”, “D. C.”, “Loew Coll.”, “Type 13428” (red label), “ PARALECTOTYPUS ♀ Anthophilina variegata Loew, J. Roháček & K. N. Barber des. 2013” (red) and “ Anthomyza variegata (Loew) ♀, J. Roháček & K. N. Barber det. 2013” (MCZC, intact).According to the original description (LOEW 1863), these specimens originate from USA: District [of] Columbia and were collected by Osten Sacken. Both type specimens belong to the pale form (see below). The lectotype and paralectotype are designated because LOEW (1863) did not explicitly enumerate specimens he used for the description and owing to the necessity to nx the status of the species inasmuch as there are another three closely related Nearctic species hitherto confused under the name A. variegata. Other material examined. CANADA: ONTARIO: Burlington, edge Tuck Ck, 43°21.2'N 79°46.6'W, pooter, Bromus inermis, 16.vii.1997, 1 ♀, sweeps/pooter, Bromus inermis, 18.vii.1997, 1 ♂, 20.vii.1997, 1 ♀; Burlington, Royal Botanical Gdn., 43°17.5'N 79°52.4'W, sweeps/pooter, Calamagrostis canadensis, 16.vii.1997, 1 ♀; same locality but 43°17.8'N 79°52.6'W, sweeps, trailside vegetation in mixed hardwood, 14.vii.2002, 1 ♂ 2 ♀♀ (all CNCI), 15.vii.2002, 3 ♂♂ 4 ♀♀ (AMNH), 16.vii.2002, 2 ♂♂ 4 ♀♀ (CASC), all K. N. Barber leg.; same locality but trailside sweeps, mostly Carex, Fragaria, Solidago, 18.vii.2002, 5 ♂♂ 5 ♀♀ (CNCI 2 ♂♂ 2 ♀♀, SMOC 3 ♂♂ 3 ♀♀, 1 ♀ genit. prep.); same locality but 43°17.79'N 79°52.61'W, trailside sweeps, mostly Carex, Fragaria, Solidago, 27.vii.2003, 2 ♂♂ 11 ♀♀ (CNCI 1 ♂ 10 ♀♀, SMOC 1 ♂ 1 ♀, both genit. prep.), sweeps, trailside vegetation, 18.viii.2014, 1 ♀ (CNCI); same locality but 43°17.78'N 79°52.61'W, sweeps, trailside vegetation in mixed hardwood, 6.ix.2005, 1 ♂ 1 ♀ (CNCI), all K. N. Barber leg.; Fergus, yard near Grand R., 27.vii.1995, 1 ♂, S. A. Marshall leg. (DEBU); Fergus, 43°41.5'N 80°23.2'W, Grand R. noodplain, sweeps/pooter, Bromus inermis, 17.vii.1997, 1 ♀, K. N. Barber leg. (DEBU); Fergus nr. Guelph, Grand River, riverside vegetation, 30.vii.1994, 2 ♂♂ 1 ♀, J. Roháček leg. (SMOC, 1 ♀ genit prep.); Lake Simcoe, Sibbald Point Prov. Pk., sweep humid forest along stream, 29.vi.2001, 1 ♂, S. E. Brooks leg. (LEMQ 0040107); Guelph, 3.vii.1979, 1 ♂, K. N. Barber leg. (DEBU); Ottawa, 19.vii.1946, 1 ♂, A. R. Brooks leg., 6.vii.1963, 1 ♂ 3 ♀♀, 13.vii.1963, 2 ♂♂ 2 ♀♀, J. R. Vockeroth leg. (CNCI); Pelee Island, Stone Rd., behind alvar, 41°45.2'N 82°38.3'W, sweeps, graminoids/ Impatiens under canopy, 8.vi.2002, 4 ♀♀, sweeps, Carex under canopy, 9.vi.2002, 2 ♂♂ 6 ♀♀, K. N. Barber leg. (DEBU); Haldimand-Norfolk Co., Port Ryerse, along stream, 15.vii.1986, 1 ♂ (headless), R. Danielsson leg. (MZLU); Rondeau P. Pk., 1.ix.1979, 3 ♂♂ 1 ♀, L. Masner leg. (DEBU); same locality but South Point Trail, 26.vi.1985, 1 ♀ (DEBU); same locality but South Point Trail, east parking lot, 42°16'48"N 81°51'17"W, sweeps, mostly Impatiens / Carex, 29.vii.2003, 4 ♂♂ 4 ♀♀ (DEBU 01500150–57); same locality but Spicebush Trail, 42°18'08"N 81°51'13"W, sweeps, mostly Impatiens / Carex, 29.vii.2003, 3 ♂♂ 3 ♀♀ (DEBU 01500142–47); Short Hills P. Pk., Terrace Ck. Trail east loop, 43°05.8'N 79°16.1'W, mostly Carex / Impatiens in mixed hardwood, sweeps, 17.vii.2002, 4 ♂♂ 1 ♀, all K. N. Barber leg. (DEBU); Simcoe, 26.vi.1939, 1 ♀, G. E. Shewell leg. (CNCI); Essex Co., Windsor, Broadway Park & Black Oak Savannah Park, 5–7.vii.2007, 1 ♀, S. M.Paiero leg.(DEBU 00299926). QUEBEC: Mt-St-Hilaire, 45°32.3'N 73°09.4'W, el. 150–200 m, sweep in deciduous forest, CJB07-02, 19.vi.2007, 1 ♂, C. Borkent leg. (LEMQ 0040536); MontSt-Hilaire Reserve, Pain de Sucre Trail, sweep along trail in forest, 27.vi.2001, 2 ♀♀, S. E. Brooks leg. (LEMQ 0040108, -09); Mont-St-Hilaire Biosphere Reserve, Pain de Sucre Trail, 27.vi.2001, sweep trail edge, 1 ♂, S. E. Brooks leg. (LEMQ 0040315), sweep at brook, 27.vi.2001, 1 ♂ 3 ♀♀, M. Pollet leg. (LEMQ 0040365, -66, -73, -76); Old Chelsea, Summit King Mt., 1150', 16.vi.1963, 1 ♀, J. R. Vockeroth leg. (CNCI); Trois Rivières, western outskirts, forest, swept, eclector, (Universität Bielefeld, X989), 25.viii.1994, 1 ♂ 1 ♀, M.v. Tschirnhaus leg. (ZSMC, in ethanol). UNITED STATES OF AMERICA: ALABAMA: Monroe Co., 1 mi S Claiborne Dam, 31°35'30"N 87°32'12"W, 29–30.v.1995, 1 ♂, J. A. MacGown leg. (MEMU); DeKalb Co., DeSoto St. Pk., 1600–1700', T6S, R10E, Sec.19W, mercury vapor & black light trap, William H. Cross Expedition, 18.v.1990, 1 ♀, D. Hildebrand & T. Schiefer leg. (MEMU). COLORADO: Evergreen, grassland slope, 22.viii.1943, 1 ♀, M. T. James leg. (USNM, genit. prep.). CONNECTICUT: Redding, 3.vi.1933, 1 ♀, A. L. Melander leg. (USNM). DISTRICT OF COLUMBIA: [no other data], 1 ♂, J. M. Aldrich leg. (USNM); Deanwood, nr. brook, 9.vi.1991, 5 ♂♂ 7 ♀♀, M. Barták leg. (MBPC 3 ♂♂ 4 ♀♀, 1 ♂ 1 ♀ genit. prep., SMOC 2 ♂♂ 3 ♀♀, 1 ♂ genit. prep.); Rock Creek, 3.viii.1913, 1 ♀, R. C. Shannon leg.; Theodore Roosevelt Island, 4.vi.1977, 1 ♀, W. N. Mathis leg.; Washington, 17.viii.1913, 3 ♂♂, A. L. Melander leg. (all USNM). GEORGIA: Lumpkin Co., 15 mi NW Dahlonega, 2.vi.1971, 1 ♀, G. Hicks leg. (LACM); Rabun Co., Pine Mountain, 1400', 4.v.1957, 2 ♂♂ 3 ♀♀, 14.v.1957, 1 ♀, J. R. Vockeroth leg. (CNCI). ILLINOIS: Champaign Co., 19.vi.1926, 1 ♀, V. Q. Smith leg. (CNCI); Williamson Co., 3 mi S Crab Orchard Lake, 17.viii.1969, 1 ♂, J. C. Marlin leg. (INHS 40,202); Dongola, 10.v.1916, 2 ♂♂ 3 ♀♀, [no collector] (INHS 40,169–73); McLean Co., Downs, 10.vi.1969, 1 ♂, D. W. Webb & J. C. Marlin leg. (INHS 40,212); Dubois, 22.v.1917, 2 ♂♂ 1 ♀, [no collector] (INHS 40,166–68); Alexander Co., Horseshoe Lake, T16S, R2N, Sec.16, sweeps, (#83032), 7.vi.1983, 1 ♂ 1 ♀, I. S. Askevold leg. (DEBU); Union Co., 0.7 mi NE Lick Creek, 24.v.1976, 2 ♂♂ 1 ♀, D. W. Webb leg. (INHS 40,203–05); Macomb, river-woods, 29.v.1962, 2 ♂♂ 1 ♀, W. W. Wirth leg. (USNM); Urbana, university forest, 28.viii.1915, 1 ♂, [no collector] (INHS 40,162); White Heath, 20.v.1916, 1 ♀, 3.vi.1917, 1 ♂, [no collector] (INHS 40,163, -64). INDIANA: Lafayette, 19.vi.1922, 2 ♂♂ 1 ♀, 22.vi.1922, 1 ♀, E. W. Stafford leg., 4.viii.1922, 1 ♀, [no collector] (MEMU); Lafayette, “viii-5”, 1 ♂, 27.vi.1916, 1 ♀ (headless), Elymus canadensis, 30.v.1917, 1 ♀ (genit. prep.), J. M. Aldrich leg. (USNM); Lafayette, 19.vii.1916, 1 ♂, [-]. vii.1931, 1 ♀, J. M. Aldrich leg. (SMOC, ♀ genit. prep.). IOWA: Ames, 25.vi.1947, 1 ♀, A. R. Brooks leg (CNCI). KENTUCKY: Franklin Co., Cove Springs Park, 38°13'N 84°51'W, 26.v.–28.vii.2005, Malaise trap, 1 ♀, Lindsay & Edelen leg. (LACM); Mammoth Cave, Nat. Pk., 23.viii.1981, 1 ♂, J. R. Vockeroth leg. (CNCI). MARYLAND: Prince Georges Co., Beltsville, 14.v.1979, 2 ♂♂ 1 ♀, A. Freidberg leg. (TAUI, 1 ♀ headless); Mongomery Co., Bethesda, 18.v.1968, 1 ♂, 26.v.1968, 1 ♂ 1 ♀, 1.vi.1968, 2 ♂♂, 2.vi.1968, 2 ♂♂ 1 ♀, 4.vi.1968, 2 ♂♂ (1 ♂ genit. prep.), 17.v.1969, 1 ♀, 25.v.1970, 1 ♀, 27.v.1972, 1 ♂, 27.vii.1972, 2 ♀♀, 29.v.1977, 2 ♂♂, 11.vi.1978, 1 ♂, G. Steyskal leg.; P[rince] G[eorge’s] Co., Camp Springs, blacklight trap, 9.vi.1979, 1 ♂, G. F. Hevel leg.; Chesapeake Beach, 30.v.1924, 1 ♀, J. R. Malloch leg.; Colesville, 11.vii.1974, 1 ♂, 4.vi.1977, 1 ♂, 14.vi.1977, 1 ♂, 18.vi.1977, 1 ♂; Montgomery Co., Colesville, Malaise trap, 26.vi.1977, 1 ♂, 3.viii.1979, 1 ♂, all W. W. Wirth leg. (all USNM); Howard Co., Dayton, Pig Tail Recreation Area, 39°13'08.37"N 77°00'23.20"W, damp undergrowth, 22.vi.2014, 6 ♂♂ 1 ♀ (GAFC 3 ♂♂ 1 ♀, 2 ♂♂ genit. prep., SMOC 3 ♂♂, 1 ♂ genit. prep., 2 ♂♂ used for molecular analysis), 20.vii.2014, 3 ♂♂ 1 ♀, G. A. & A. M. Foster leg. (GAFC); Montgomery Co., Dickerson, 14.vii.1974, 1 ♂ 1 ♀, G. A. Foster leg.; College Park, 7.v.1969, 1 ♂, G. F. Hevel leg.; Glen Echo, 21.viii.1921, 1 ♀, 16.vii.1922, 1 ♀, 20.v.1923, 1 ♂, J. R. Malloch leg.; Lavale, 9.v.1970, 1 ♀, G. Steyskal leg. (all USNM); Prince Georges Co., Patuxent Nat. Wildlife Refuge, 30.vi.1986, 1 ♂, R. Danielsson leg. (MZLU, genit. prep.); Plummers Island, 9.vi.1913, 1 ♀, R. C. Shannon leg. (SMOC); Plummers Island, 23.vi.1907, 1 ♂ 1 ♀, 2.vi.1912, 1 ♀, 9.v.1913, 1 ♂, 17.vi.1913, 2 ♀♀, 3.viii.1913, 1 ♀ (headless), 7.vi.1914, 1 ♀, W. L. McAtee leg., 5.viii.1913, 1 ♀, 8.v.1915, 1 ♂ 1 ♀, R. C. Shannon leg., 29.viii.1971, 1 ♀, G. F. Hevel leg.; near Plummers Island, at light, 12.viii.1914, 1 ♀, R. C. Shannon leg.; Somerset Co., Snow Hill, swamp margin, 19.v.1968, 1 ♀,W.W.Wirth leg. (all USNM); Montgomery Co., Wheaton, 27–28.v.1979, 1 ♂, 25.vi.1979, 1 ♂ 3 ♀♀, 3.vii.1979, 1 ♀, 16.viii.1979, 1 ♂ (headless) 1 ♀ (genit. prep.), A. Freidberg leg. (TAUI). MASSACHUSETTS: Catoctin, Mt. Park, Chestnut wood, 15.vi.1991, 2 ♂♂ 2 ♀♀ (MBPC, 1 ♂ genit. prep.); Catoctin, Mt. Park (Lantz), 15.vi.1991, edge of wood, 1 ♂ (MBPC), meadow nr. pond, 1 ♂ 1 ♀ (SMOC 1 ♂, MBPC 1 ♀, genit. prep.); Catoctin, Mt. Park, Owen’s Creek, 15.vi.1991, 3 ♀♀, all M. Barták leg. (MBPC 2 ♀♀, SMOC 1 ♀, genit. prep); Concord, fern bog, 22.vii.1961, 1 ♀, W. W. Wirth leg. (USNM); Franklin Co., ~0.5 km E Farley, 42°36.16'N 72°25.94'W, sweeps, asters, ferns, Impatiens, Rubus, under canopy, 26.vii.2006, 1 ♀, K. N. Barber leg. (CNCI); Petersham, [-]. vii.1926, 1 ♂ 1 ♀ (♂ headless), 30.vii.1926, 1 ♂, A. L. Melander leg. (USNM); Woods Hole, 20–30.vi.1913, 1 ♂, 13.vii.1913, 1 ♂ (USNM)

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