Crassomicrodus fulvescens Cresson 1865

Crassomicrodus fulvescens (Cresson 1865) (Fig. 1 a–f) Microdus fulvescens Cresson 1865: 297 [Examined]. Microdus medius Cresson 1865: 298 [Examined]. New synonymy. Holotype: Female. Col. No. 1727.1 (ANSP). Holotype of M. medius: Male. Col. No. 1725 (ANSP). Description: Female. Color: Integument yellowish orange except eye silver (Fig. 1 a, b) or blackish, ocelli translucent yellow (Fig. 1 d); antenna black; apical area of hind tibia and tarsomeres blackish, sometimes apical area of middle tibia, tarsomeres and/or trochanter blackish; wing veins dark brown; forewing infumate with large hyaline spot in first submarginal and second discal cells. Head: Transverse in frontal view; face with weak longitudinal ridge dorsomesally; eye height/width = 1.48–1.55; eye height (lateral view) 0.59–0.61X inter-ocular distance (anterior view); pyramidal-shaped raised surface between antennae with two weakly defined tubercles (Fig. 1 b, c); frons deeply excavated; posterior area of antennal sockets smooth (Fig. 1 d); groove between lateral ocelli smooth; median ocellus separated from lateral ocellus by smooth groove; gena distinctly bulging (Fig.1 b, c); malar space (anterolateral view) 0.55–0.65X longer than eye height; clypeus (anterior view) 2.44–2.67X wider than high; length of ventrolateral margin of clypeus similar to diameter of tentorial pit; antenna with 34–37 flagellomeres; setae at base of mandible distinctly longer than setae on rest of body surface. Mesosoma: Pronotum smooth (Fig. 1 e); lateral pronotal margins with superficially crenulate groove; notauli impressed; anterolateral edges of scutellum lacking small acute projection (Fig. 1 f); scutellar disc convex with sparse setae from 0.08–0.09 mm in length; scutellar disc sloped posteriorly and rounded; lateral scutellar depression rugose and foveolate (Fig. 1 f); carinae of central metanotal area almost circular shaped; propodeum reticulate rugose; subalar lobe separated from mesopleuron by narrow rugulose groove, with rugosities posteriorly (Fig. 1 e); metapleuron reticulate-rugose. Legs: inner spur of middle tibia 0.54–0.58X length of basitarsus; inner spur of hind tibia 0.48– 0.54X length of basitarsus; metabasitarsus 1.10–1.24X length of tarsomeres III, IV, and V combined; hind tibia 2.17–2.38X longer than basitarsus; hind femur length 3.70–4.00X its maximum width. Wings: forewing length/width = 2.61–2.72; stigma 3.38–3.85X longer than maximum width; forewing vein R1 0.56–0.66X as long as vein RS; vein RS not sinuate; vein r arising beyond middle of stigma; second submarginal cell triangular, with petiole 0.13–0.24 mm long; vein M+CU distinctly pigmented throughout; hind wing length/width = 3.46–3.56; hind wing vein 1M 1.47–1.71X longer than 1r-m; hind wing with 5–7 hamuli. Metasoma: Apical width of petiole (tergum 1) 2.73–2.97X wider than basal width (dorsal view); minimum width of petiole 0.61– 0.62X apical width (dorsal view); length of ovipositor sheath 0.24–0.33 mm. Body length: 6.35–8.05 mm. Male: Similar to female except color as follows: integument black except medial area of mandible, pronotum, mesonotum, subalar lobe, tegula, metasoma, femora, tibia, and tarsomeres yellowish orange. Apical area of hind tibia and all hind tarsomeres sometimes blackish. Host: The alfalfa looper, Autographa californica (Speyer) (Lepidoptera: Noctuidae), is reported as a possible host for C. fulvescens based exclusively on label data from a female specimen in the USNM from Walla Walla, Washington. However, isolated rearings are necessary to verify A. californica as a host since host remains are not associated with the specimen. Autographa californica is distributed throughout the United States and parts of Canada. Larval host plants include pea, sugarbeet, alfalfa, bean, mint, and spinach (Berry 1998). Specimens examined: Holotype Ψ M. fulvescens: Col. (ANSP). Holotype ɗ M. medius: Col. (ANSP). Other specimens examined: CANADA, British Columbia: Osoyoos: 1 Ψ 4 ɗ 21/VII/1953, McGillis J.R. (1 Ψ 3 ɗ CNC, 1 ɗ HIIC); 1 ɗ 22/VII/1953, McGillis J.R. (CNC); 4 ɗ (1 homotype) 25/VIII/1953, Martin J.E.H. (3 CNC, 1 HIIC); 1 Ψ 3 ɗ 27/VII/1953, Martin J.E.H. (1 Ψ 2ɗ CNC, 1 ɗ HIIC). MEXICO, Chihuahua: Boquilla, 1 Ψ 23/VII/1987, González A. (CIBE-UANL). Coahuila: 39 km S Agua Nueva, 1 ɗ 20/X/1994, 1770 m., Mercado I., 24.53.21 N 101.04.63 W (TAMU). Sonora: Minas Nuevas, 1 ɗ 7/VIII/1952, Vaurie C. & Vaurie P. (AMNH). Zacatecas: 9 miles S Fresnillo, 2 Ψ 9/VII/1954, Linsley E.G, MacSwain J.W. & Smith R.F. (EMEC, USNM). USA: Ckll, 1 ɗ 4523, 1926; 1 Ψ [no date]; 1 Ψ 17/VII/1877, Greeley; Pasco, 4 ɗ 26/V/ 1894, Piper C.V. (1 AEIC, 3 USNM). Arizona: Cochise Co.: Portal, 1 Ψ 2–12/IX/1976, Van der Vecht J. (AEIC). 2 miles E Portal, 1 Ψ 25/VIII/1966, Gertsch W.J.; 2 miles NE Portal, 1 ɗ 1/IX/1960, Cazier & Feight; 2.5 miles NE Portal, 1 ɗ 26/VIII/1959, Statham M.; Skeleton Cyn., Peloncillo Mountains, 1 Ψ 4/V/1966, Rozen & Favreau (AMNH). 1.5 miles W St. David, 1 Ψ 3/IX/1961, Hurd P.D. (EMEC). Chiricahua Mountains, Portal: 1 Ψ 1 ɗ 15/VIII/1958, Bohart R.M.; 1 Ψ 6/VIII/1958, James R.H.; Portal: 1 Ψ 16/VIII/1958, Marsh P.M.; 3 Ψ 1 ɗ 16/VIII/1958, Moore C.G.; 1 Ψ 5/VIII/1958, Bohart R.M.; 85 miles W Tombstone, 1 ɗ 1/ VIII/1966, Kovacic C.R.; San Simon, 1 ɗ 13/VIII/1981, Bohart R.M.; Willcox, 1 Ψ 28/VIII/1974, Bohart R.M.; 7 miles N Tucson, Pima Co., 1 Ψ 4/IX/1968, Miller D.R. & Lauck J.E. (UCDC). Tucson, 1 ɗ 6/IX/ 1933, Bryant (CAS). N Sonoita, 1 ɗ 10/VIII/1959 (CNC). Continental, 1 Ψ 24/IV/1958, Butler G.D.; Portal Creek Cyn., 1 ɗ 5/VIII/1977, Masner L. (HIIC). Vernon, 1 Ψ 25/VI/1957, Butler G. & Werner F. (USNM). Arkansas: Willcox, 1 Ψ 1 ɗ 31/VII/1909, Fisher A.K. (USNM). California: Oak Glen, San Bernardino Co.: 1 Ψ 25/VII/1984, 1500 m., Wagner Robert E., Malaise trap; 4 Ψ 31- VIII/6 - IX/1984, 1500 m., Wagner Robert E.; 1 ɗ 12–18/X/1984, 1500 m., Newton and Thayer (CNC). 18 miles E Camp Ozena, Ventura Co., 1 Ψ 2/VII/ 1965, Powell J.; Sand dunes, 1 mile S Rio Vista, Solano Co., 2 Ψ 7/VIII/1976, Whitman Doug (EMEC). Menifee Valley, Riverside Co., 1 Ψ 20/VIII/1976, Frommer S.I. & Frommer S.L., 117.12.45 N 33.39.19 W (FSCA). Reedley, Fresno Co., 2 Ψ 3 ɗ 7– 10 /VII (UCDC). Valley Field Station, UC Moreno, Riverside Co., 1 Ψ 10/V/1979, (UCR). Madera, Madera Co., 1 Ψ 5/X/1975, Linsley E.G., J.M. & Michelbacher A.E., M.M. (USNM). Colorado: La Junta, 2 Ψ 2 ɗ 12/VIII/1920, 1250 m., 37.59 N 103.31 W; Pueblo, 1 Ψ 9/VIII/1920, 1433 m., 38.10 N 104.36 W (AMNH). Baca Co., 1 Ψ VI/1939, Landburg R. (CAS). Rocky Ford, 1 Ψ 26/VI/ 1917, Popenoe C.H.; Estes Park, 1 Ψ (homotype) 2/VII/1961, 2286 m., Mason W.R.M. (CNC). Wray, 1 Ψ 17– 19/VIII/1919, 40.0 N 102.10 W, 3,700 ft (HIIC). Colorado, 1 Ψ VII/1916, Popenoe; Hartman, 1 Ψ 14/VII/ 1955, Marston Norman; Hartman, 1 ɗ 21/VI/1957, Marsron N. (KSUC). Chimney, 1 ɗ (MCZ). Pueblo, 1 Ψ 11/VI/1956, Dreisbach R. & Dreisbach K. (MSUC). Inspiration, Denver, 1 ɗ 2/VII/1933, Gibbons H.I. (UCMC). Canon City, 1 Ψ; Colorado, 2 Ψ 1 ɗ; 0.5 miles SE Earl, 1 ɗ 23/VIII/1929, Romey V.E.; Rocky Ford: 1 Ψ 10/VII/1954, Titus E.S.G.; 4 ɗ 23VII/16 VIII/1909, Marsh H.O.; 1 Ψ 26/VI/1917, Popenoe C.H.; 1 ɗ 5/IX/1909, Marsh H.O.; 1 ɗ 9/VII/1912, Marsh H.O. (USNM). Idaho: Montpelier, 1 Ψ 2 ɗ 6/VII/1920, 1859 m., 42.19 N 111.18 W (AMNH). 1 Ψ 2 ɗ Preston, 17/VII/1922, Van Duzee E.P. (CAS). Burley, 2 Ψ 27/VI/ 1932, Fox David E.; Hansen, 1 ɗ 29/VII/1932, Fox David E.; Oakley: 2 ɗ 1/VII/1927, Carter W.; 1 ɗ 14/VII/ 1927, Carter W.; 1 ɗ 17/VIII/1927, Carter W.; 1 Ψ 19/VI/1933, Fox David E.; 2 ɗ 25/VI/1928; 3 ɗ 29/VII/ 1932, Fox David E.; 1 ɗ 30/VIII/1932, Fox David E.; Paul, 1 Ψ 20/VI/1930 (USNM). Kansas: Seward Co., 1 ɗ 27/VI, Lantz, (KSUC). Garden City, 2 Ψ VI/1896, Menke H.; Garden City, 1 Ψ VIII/1896, Menke H. (USNM). New Mexico: Hatch: 1 ɗ 28/VIII/1974, Townes H. & Townes M.; 2 Ψ 1 ɗ 29/VIII/1974, Townes H. & Townes M.; 1 ɗ 30/VIII/1974, Townes H. & Townes M.; Nutt, 1 Ψ 29/VIII/1974, Townes H. & Townes M.; Rodeo, 1 Ψ 7/IX/1974, Townes H. & Townes M.; Tucumcari, 2 ɗ 13/V/1981, Dasch C. & Dasch B. (AEIC). Cubero, Valencia Co., 1 ɗ 18/VIII/1948, Vaurie C. & Vaurie P. (AMNH). Santa Fe: 1 ɗ 22/VII/1926, Van Dyke E.C.; 1 Ψ 24/VII/1926, Van Dyke E.C. (CAS). 25 miles N Las Cruces, Dona Ana Co., 1 ɗ 24/IX/1974, Bohart G. & Hanson W. (CNC). 7 miles NE Albuquerque, Bernalillo Co., 1 Ψ 1 ɗ VII/1955 (EMEC). Oasis St. Park, Roosevelt Co., 1 ɗ 31/VIII/1971, 1250 m., Brown K.W. & Petrulis P.J. (ESUW). 2 miles NW Rodeo, Hidalgo Co., 1 Ψ 31/VIII/1960, Cazier & Feight (HIIC). 16 miles S Datil, Catron Co, 1 Ψ 16/VI/1956, Dreisbach R. & Dreisbach K.; Hoxie Jc. [Colfax Co.], 1 Ψ 2 ɗ 12/VI/1956, Dreisbach R. & Dreisbach K.; Rotan, 1 Ψ 12/VI/1956, Dreisbach R. & Dreisbach K. (MSUC). 15 miles N Rodeo La Cienega L., Hidalgo Co., 1 ɗ 10/ IX/1971, Villegas B.; Carrizozo, Lincoln Co., 1 Ψ 6/VIII/1966, Kovacic C.R.; Rodeo, Hidalgo Co.: 1 Ψ 1 ɗ 10/VIII/1958, Rice R.E.; 1 ɗ 2/VIII/1958, Rice R.E.; Tucumcari, Quay Co.: 1 ɗ 10/VII/1963, Bohart R.M.; 2 Ψ 7 ɗ 17/VIII/1967, Bohart R.M. (UCDC). 14 miles SW of Portales, nr. Boone Draw, Roosevelt Co., 4 Ψ 26/ V/1972, Frommer Saul & Jorgrnsen N. (UCR). 0.5 miles W Springer, 1 Ψ 23/VIII/1929, Romney V.E.; 2 miles N San Jon, 1 ɗ 12/VII/1929, Romney V.E.; 16 miles S Datil, Catron Co., 1 ɗ 16/VI/1956, Dreisbach R. & Dreisbach K.; 5.7 miles W Grady, 1 ɗ 28/VI/1929, Romney V.E.; Correo, Valerica Co., 1 Ψ 15/VI/1956, Dreisbach R. & Dreisbach K.; Jemez Springs, 1 ɗ 21/VII/1929, Englehardt G.P.; Hoxie Jc. [Colfax Co.], 1 Ψ 1 ɗ 12/VI/1956, Dreisbach R. & Dreisbach K.; Las Cruces, Cockerell 1 Ψ; Las Cruces, 1 ɗ V/1923?, Garcia (USNM). Oklahoma: Forgan, 1 Ψ 1 ɗ 14/VI/1935, Brues (MCZ). Oregon: Hermiston, 2 ɗ 9/VII/1922, Melander A.L. (MCZ). Hermiston, 1 Ψ 10/V/1930, Scullen H.A. (USNM). Texas: Davis Mountains, 1 Ψ 11/ VIII/1957, Arnett R.H. (CNC). 20 miles S Kent, Jeff Davis Co., 1 Ψ 15/VIII/1965, Schaffner J.C. (HIIC). Conlen, 1 Ψ 7/VIII/1952, Dreisbach R.R.; El Paso, 1 Ψ 20/VI/1909, Bishop F.C. (U

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