Quedius levicollis

Quedius levicollis (Brullé, 1832) (Figs 1, 2B, 3, 6D, 7D, 11B, 19) Staphylinus levicollis Brullé, 1832: 131 [Type locality: Morée: plaine de Modon] Philonthus frontalis Nordmann, 1837: 76 [Type locality: Gallia; Symperopoli] Quedius balthasari Mařan, 1935: 9 [Type locality: Knossos, Creta] Staphylinus nigritarsis Ménétriés, 1832: 144 [Type locality: Caucase] References. GඋൺඏൾඇHඈඋඌඍ (1802):34,(1847):231 (characters); SඍൾඉHൾඇඌ (1832): 215, (1839): 389 (characters); Fൺඅൽൾඋආൺඇඇ (1835): 123 (characters); Lൺർඈඋൽൺංඋൾ (1835): 376 (characters); Nඈඋൽආൺඇඇ (1837): 75 (distribution); Hඈඅආൾ (1837): 63 (distribution); Wൾඌඍඐඈඈൽ (1838): 167 (larva and pupa); EඋංർHඌඈඇ (1840): 537 (characters); Kඈඅൾඇൺඍං (1846): 22 (characters); Kංൾඌൾඇඐൾඍඍൾඋ (1848): 54 (characters); HඈർHHඎඍH (1849): 154, (1862): 46 (characters); RൾൽඍൾඇൻൺർHൾඋ (1849): 709, (1857): 201, (1874): 198 (characters); Fൺංඋආൺංඋൾ & Lൺൻඈඎඅൻජඇൾ (1856): 536 (characters); RඈඌൾඇHൺඎൾඋ (1856): 71 (distribution); Kඋൺൺඍඓ (1857): 502 (characters); Fൺංඋආൺංඋൾ & Cඈඊඎൾඋൾඅ (1860): 154 (distribution); Fൺඎඏൾඅ (1865): 293 (ecology); (1874): 289 (characters and distribution); (1878): 128, (1886): 58, (1902): 115 (distribution); Mඎඅඌൺඇඍ & Rൾඒ (1876): 688 (characters); Fඈඐඅൾඋ (1888): 237 (characters); GൺඇGඅൻൺඎൾඋ (1895): 403 (characters); Sൺංඇඍൾ- Cඅൺංඋൾ Dൾඏංඅඅൾ (1906): 102 (distribution); Pඈඋඍൺ (1907): 129 (characters); Rൾංඍඍൾඋ (1909): 111 (characters); JඈHൺඇඌൾඇ (1914): 363 (characters); Mඎඇඌඍൾඋ (1923): 196 (distribution); Gඋංൽൾඅඅං (1924):80 (characters and distribution); Pඈඋඍൾඏංඇ (1929):341 (characters); WඳඌඍHඈൿൿ (1938): fig. 22 (characters); TඈඍඍൾඇHൺආ (1949): 376 (characters); Hൺඋൽඒ (1851): 32 (distribution); Hൺඇඌൾඇ (1952): 142 (characters); SർHൾൾඋඉൾඅඍඓ (1956): 1102, (1965a): 407, (1965b): 511 (distribution); (1961): 133 (characters); Sආൾඍൺඇൺ (1958): 362 (characters and ecology); (1962): 134 (characters and distribution); (1963): 39, (1967): 558, (1970): 62, (1993):50 (distribution);(1978b):121 (distribution and ecology); Pൺඅආ (1963): 141 (characters and distribution); LඈHඌൾ (1964): 211 (characters); Fඈർൺඋංඅൾ (1964): 61 (distribution); Hඈඋංඈඇ (1965): 273 (distribution); Cඈංൿൿൺංඍ (1970):62 (distribution); (1978):192 (characters and distribution); Bඈඋൽඈඇං (1976a):93 (characters);(1982):140 (ecology); (1984):342 (distribution); Pඈඉൾ (1977): 31 (characters and distribution); Hංඇඍඈඇ (1981): 662 (characters of egg); TඬඍH (1984): 118 (distribution); Fඋൺඇඓ (1987): 70 (distribution); Gඎඌൺඋඈඏ (1993): 73 (synonymy); WൾඅർH (1993): 229 (ovary structure); Cංർൾඋඈඇං & Zൺඇൾඍඍං (1995): 33 (distribution); Nൾඐඍඈඇ (1995): 51 (requested to ICZN to place Q. levicollis on the Official List of Specific Names in Zoology); ICZN (1996): 215 (Q. levicollis placed on Official List of Specific Names in Zoology); Bඅൺඇൽ & Sංඇർඅൺංඋ (1997): 204 (biology); MൺඋർHൺඅ & CH ൺඋൽඈඇඇൾඍ (2000): 72 (distribution); Oඐൾඇ (2000): 252 (ecology); AඌඌංඇG & Wඎඇൽൾඋඅൾ (2001): 37 (distribution); UHඅංG et al. (2006): 54 (distribution); Tඋඈඇඊඎൾඍ (2006): 102 (distribution); Aඇඅൺ෡ (2009): 229 (distribution); Sඍൺඇ (2009): 242 (distribution); Vඈඋඌඍ et al. (2009): 16 (distribution); Mൺඃඓඅൺඇ & BඈHගඹ (2012): 182 (distribution); Bൾൻൾඋආൺඇඌ et al. (2016): 135 (distribution); ÖඓGൾඇ et al. (2016): 620 (distribution); Rඎඍൺ et al. (2016): 37 (distribution); Sൺඅඇංඍඌĸൺ & Sඈඅඈൽඈඏඇංĸඈඏ (2019): 49 (distribution). Type material examined. Staphylinus nigritarsis: Lൾർඍඈඍඒඉൾ: designated by Gඎඌൺඋඈඏ (1993) (ZIN), labelled: “[golden square] / Caucas. / nigritarsis Menet. Cauc ”. Pൺඋൺඅൾർඍඈඍඒඉൾ: labelled: “[golden square] / Caucas.” (1 J ZIN). Additional material examined. ALBANIA: 13km SE Elbasan, Mali i Shpatit, Mt. Bukanikut, 41.0431, 20.2172, brook bank in beech forest, 1500m, 22.V.2010, leg. D.W.Wrase (1J cSch). BOSNIAAND HERZE-GOVINA: Maklen-Pass,[43.85,17.65], 1902, leg.O.Leonhard (1 SDEI); Prozor, [43.82, 17.61], 1902, leg. O. Leonhard (1 SDEI). BULGARIA: Arkutino, S of Burgas, [42.41, 27.65], 28.V-9.VI.1971, leg. Wallis (1 J SDEI). CROATIA: Capela [Kapela], [44.89, 15.52], leg. Heyden (1 SDEI); Velebit Mountains, [44.52, 15.22], wine traps, V.2005 (1 J cJen). CYPRUS: Agios Nikolaios, [34.86, 32.75], 16.IV.1995, leg. H. Schmid (1 J NMW); Troodos Mountain, Stavros tis Psokas, [35.02, 32.63], 800- 950m, 12.-18.IV.2010, leg. I. Wolf (1 J cSch). DENMARK: Ovestrup Hede, 56.2412, 8.9297, pitfall trap, heathland grazed part, 7-22.VI.2017, leg. E. Kristensen, D. Byriel, & M. Justesen (1 J NHMD). FRANCE: Avignon, [43.94, 4.81], 22.V.1949, leg. R. Skovgaard (1 J NHMD); Chigny n. Oise, [49.91, 3.76], 19.X.1918, leg. Salchert (1 ♀ ZMHB); Paris, [48.83, 2.41], 19.V.1949, leg. R. Skovgaard (1 J NHMD). Cඈඋඌൾ: Bocognano,[42.08,9.06],1905, leg.O.Leonhard (1 SDEI);Col de Bavella, [41.79, 9.23], 1240m, East slope, 25.VII.1994 leg. L. Zerche (1 J SDEI); Foret de Valdo-Niello, Evisa, [42.28, 8.91], 1000-1200 m, 27.VII.1967, leg. Langemark-Lomholdt (1 J NHMD). GERMANY: Berlin-Marzahn, Hellersdorfer Berg, [52.53, 13.58], 8.VIII.1997, leg. Hieke (1 ♀ ZMHB); Berlin, Prenzlauerberg Hof, [52.53, 13.46], 29.V.1985, leg. I. Uhlig (1 J ZMHB); Grunewald n. Zehlendorf, [52.48, 13.22], leg.Wagner (1 SDEI). GREECE: Attica, [38.05, 23.84], 17.IV.1922, leg. W. Liebmann (1 SDEI); Paleokastro, Halkidiki, [40.43, 23.42], 800m, 11.VII.1983, leg.A. Casale (1 ♀ cBor); 2 km W Panagia, Trikala, [39.79, 21.33], 15.VI.1982, leg. R Danielsson (1 J MZLU); Voria Pindos, Morfi, [40.21, 21.21], dry slope, 26.VI.1997, leg. Winkelmann (1 ♀ cSch); Cඈඋൿඎ: Pandokratoras, 39.7492, 19.8701,N-slope with scattered pine,decidious trees,and bushes, litter and roots between rocks sifted, 800 m, 4.VI.2017, leg. Schülke (1 J cSch);Pantokrator,[39.74,19.87], V.1964, leg.Palm (1♀ MZLU). Cඋൾඍൾ: Georgioupouli, [35.36, 24.26], 24.-25.IV.1995, leg. B. Jaeger (1 ♀ cSch); Ida mts., ca. 6 km S Analipsis, 35.1752, 24.8391, Quercus forest, 1260m, 21.X.2006, leg. M. Schülke (1 J cSch); Mt.Ida, 28km SW Iraklio, [35.23, 24.77], 1600m, 7.IX.1991, leg. N. Esser (1 ♀ NHMD). KൺඋඉൺඍHඈඌ: Karpathos, 35.5097, 27.2189, 40 m, terracod grassy fallow, 01.I.2016, leg. V. Assing (ZMHB). Lൾඌൻඈඌ: SW Agiassos, road to Olympos peak, [39.06, 26.36], 700 m, 10.VI.1996, leg. Jäch (1 J 1 ♀ NMW). Pൾඅඈඉඈඇඇൾඌ: Korinthia, Kyllini, Zoria, [37.94, 22.43], 2000 m, 3.VI.1995, leg. Giachino & Vailati (1 J 3 ♀♀ cBor); Kyllini Mountain, [37.97, 22.39], 2100m, North side, below summit over valley, 9.VI.1996, leg. P. Wunderle (1 ♀ cSch); Sparta, [37.06, 22.43], 3.V.1962, leg. Skovgaard (1 J NHMD); N Taygetos, 6 km NE Thouria, 37.1401, 22.1284, litter and soil under bushes, sifted, 584 m, 20.IV.2015, leg. M. Schülke (1 ♀ cSch). RHඈൽൾඌ: Prophitis Ilias, 36.2734, 27.9287, pine forest, under stones & sifted from litter & moss, 685m, 17.V.2015, leg. Wrase & Laser (1 J cSch). IRAN: Kohgiluyech-Va-Boyern-Ahmad province, 20 km SW Yasuk, 10 km SE Sepidor, [30.56, 51.51], 5.-6.V.2007, leg.A. Anistchschenko (1 ♀ SDEI). IRELAND: Inch Dingle Eire, [52.14, -9.97], IX.2012, leg. J.J. Shaw (1 J cJen); Pembury, Kent 51.1625, 0.3431, 24.VII.2011, leg. J.J. Shaw (2 JJ cJen). ISRAEL: Upper Galilee,Har Avivim,W Avivim, 33.0833, 35.4923, open stony grazing land limestone, 768m, 21.IV.2006, leg. D.W.Wrase (1 J cSch); Upper Galilee, Meron Mts. Har Meron, Kamin Rom, 32.9908, 35.4111, open stony grazing land, limestone, 1100m, 28.IV.2006,leg.D.W. Wrase (1J cSch). ITALY: CalabriaAspromonte,[38.21, 15.99],1905,leg. Pagametti (1 SDEI); Calabria, Roseto Capo Spulico, [39.98, 16.61], 20 m, 6.X.1998 leg.F.Angelini(1♀ NHMD); Dicomano, Toscana,[43.89,11.52], I.1971, leg Bordoni (1J cBor); Macerata, [43.29, 13.45], 1.X.1955, leg.D. Caccamicelo (1♀ cBor); Marche,Montemonaco, Isola San Biagio,[42.91, 13.31], fields, 900 m, 27.VII.-1.IX.2004, leg.M.Rismondo & R. Fabbri (2 JJ 2♀♀ NHMD); same locality, black hornbeam forest, 1.IX.-7.X.2004, leg. M. Rismondo & R. Fabbri (1 J NHMD); Modena, Prignano sulla Secchia, La Quercia, [44.45, 10.73], pitfall traps, 400 m, 5.-18.VII.2006, leg. R. Fabbri (1 J NHMD); Reggio di Calabria, river Sant Agata, [38.09, 15.66], 11.III.1959 (1 J cBor); Settecamini, Rome, Lazio, [41.93, 12.62], 23.Xi.1969 (1 J 1 ♀ cBor); Sovara, Anghiari, [43.53, 12.03], 19.V.1930, leg. A. Andreini (1 ♀ cBor). Sൺඋൽංඇංൺ: Aritzo Mountain, Gennargentu, [40.01, 9.29], leg. C.Kruger (2 SDEI); Ottana, [40.23, 9.04], 15.X.1974 (3 JJ 1♀ cBor);Oristano,[39.89,8.59], 3.V.1975,leg.S. Ripze (2♀♀ cBor); Sංർංඅඒ: Ficuzza, [37.85, 13.38], 1906, leg. O.Leonhard (1 SDEI);Messina, [38.18, 15.54], leg. Schiödte (4 ♀♀ NHMD). MOROCCO: Ain Leuh, [33.28, -5.33], 17.V.1925, leg. M. Escalera (1 J 4 ♀♀ MNCN); Ar Rif, Chefchaounen reg., env.Bad Bessen, 34.9913, -4.8555, 1124 m, 3.VI.2007, leg. P. Hlaváč (1 J cSch); Ar Rif, Chefchaounen region, rd. Bad Berret - Bad Bessen, 1392 m, 35.0169, -4.0035, 3.-6.VI.2007, leg. P. Hlaváč (1 ♀ cSch); Taza, [34.22, -4.00], leg. F. Schubert (1 ♀ NMW). PORTUGAL: Penedo da Meditação, Coimbra, [40.224371, -8.40], 24.IV.1961, leg.R.N. F. (1 J NMW). Mൺൽൾංඋൺ: Faja da Nogueira, [32.74, -16.90], laurisilva with big Ocoteas, 750 m, 20.IX.1980, leg. H. Enghoff & O. Martin (2 JJ 3 ♀♀ NHMD); Paul da Serra, [32.73, -17.05], 27.XII.1987, leg.Gillerfors (1 J MZLU). SPAIN: Castanos de Calabazas, 1 km S Castanar de Ibor, Extemadura, 39.6235, -5.4128, 860 m, sifted mossy rock scree litter, chestnut and oak forest /w rock screes and small stream, 10.V.2019, leg.A. K. Hansen, J. J. Shaw, & J. Kypke (NHMD); Cavarrubias [Covarrubias], Burgos,[42.05, -3.51], 10.XI.1995, leg.A.Anichtchenko(1J cSha); Lloret de Mar, [41.69, 2.83], 17.IX.-6.X.1961, leg. Palm (1 J MZLU); Sierra de Ancares,A Degrada vill., [42.81, -6.87], 1100-1250 m, 8.-9.VII.1996, leg. Wrase (2 JJ cSch); Sierra de Guadarrama, Navacerrada,Arroyo de Peca [Peña] Cabrita, [40.77, -3.99], 1800 m, 5.IX.2001, leg.A.Anichtchenko (1 J NHMD); Sierra Nevada,rio Lanjaron,Tello,[36.92, -3.47], 1500 m, 29.- 31.III.2010, leg.Shavrin &Anichtchenko (1J NHMD);Toledo,El Pielago Navamorcruende, [40.15, -4.79], 4.VIII.2005 leg. Lencina & Andujar (1 ZMHB). SYRIA: Krak des Chevaliers,

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