Rhamphomyia (Dasyrhamphomyia) nigrita Zetterstedt

Rhamphomyia (Dasyrhamphomyia) nigrita Zetterstedt (Figs 14–16) Rhamphomyza nigrita Zetterstedt, 1838: 567. Type-locality: Greenland. Other references: Bezzi, 1903: 227 (catalogue); Melander, 1928: 199 (catalogue); Collin, 1931: 67 (faunal survey); Henriksen, 1939: 80 (Greenland list); Oliver, 1963: 178 (species list); Melander, 1965: 465 (catalogue); Gorodkov & Kovalev, 1969: 631 (key); Downes, 1970: 770 (phenology); Danks, 1981: 465 (arctic insects); Barták & Danielson, 2007: 107 (type data). R. (Dasyrhamphomyia) nigrita: Frey, 1922: 66 (key); Frey, 1955b: 518 (revision). R. (Rhamphomyia) nigrita: Chvála & Wagner, 1989: 288 (catalogue); Shamshev, 2001: 333 (key); Yang et al., 2007: 186 (catalogue); Barták, 2015: 576 (Greenland fauna); Shamshev, 2016: 78 (checklist). Empis borealis Fabricius, 1780: 211 (junior primary homonym of Empis borealis Linnaeus, 1758). Type-locality: Greenland. Rhamphomyia conservativa Malloch, 1919: 48. Type-locality: Bernard Harbour, Nunavut, Canada. Other references: Melander, 1928: 189 (catalogue); 1965: 462 (catalogue); Downes, 1970: 770 (as new synonym of R. nigrita); Cooper & Cumming, 1993: 32 (type catalogue). Notes on synonymy. Rhamphomyia conservativa was described from nine specimens, of which seven were found in CNC. This synonym was long recognized in the Empididae collection of the CNC and Downes (1970) clearly proposed this synonymy, which we can also confirm. This synonymy was apparently unknown to Melander (1965). Type material examined. Rhamphomyia conservativa: HOLOTYPE ♂ (Fig. 14A), labelled (Fig. 14B): “HO- LOTYPE/ Rhamphomyia / conservativa Mall. / CNC No. 219”; “W. of Bernard Isl/ N.W. T.”; “ F. Johansen / Coll.”; “Canadian/ Arctic/ Expedition/ July 14, 1916 ”; “ Rhamphomyia / conservativa/ Mall. Type” (CNC). PARATYPES: CANADA. Nunavut: Bernard Harbour, 10.vii., 1–7.viii.1915, 10, 14.vii.1916, F. Johansen (1 ♂, 4 ♀, CNC). Yukon: Herschel Island, 29.vii.1916, F. Johansen (1 ♀, CNC). Additional material examined. CANADA. British Columbia: Summit Lake, mi 392 Alaska Hwy, 5000 ft, 6.vii.1959, R.E. Leech (1 ♀, CNC); same locality, 4700 ft, 7.vii.1959 (1 ♀, CNC). Manitoba: Churchill, 14.vii.1947, C. R. Twinn (2 ♂, CNC); Churchill, 12.vii.1948, W. R. Richards (1 ♂, CNC); Churchill, 29.vi.1948, G.E. Shewell (1 ♀, CNC); Fort Churchill, 4.vii.1952, J.G. Chillcott (1 ♀, CNC); Eastern Cr. nr. Churchill, 25.vi.1952, J.G. Chillcott (1 ♀, CNC); Churchill, 10–25.vi.1930, O. Bryant (7 ♂, 4 ♀, CNC); Churchill, 58°45′16″ 93°59′55″W, A frame, MT, 26–30.vi., 30.vi.–3.vii.2007, A. Renaud (14 ♂, 13 ♀, CNC); Churchill, 58.73351°, -93.79729°, Repl.1, mesic, MT, 5–9.vii.2010, NBP field party, CCDB-21421-G08 (1 ♀, barcode associated, LEM); Churchill, Twin Lakes, MT, 58°37′53″ 93°47′20″W, 26–29.vi.2007, J. Skevington (2 ♂, 1 ♀, CNC). Northwest Territories: Aklavik, 29.vi.–18.vii.1931, O. Bryant (5 ♀, CNC); Banks Is., Aulavik NP, 73.233284°, -119.54719°, Repl.2–3, wet & mesic, Pans/ sweeps, 7-19.vii.2011, NBP field party (4 ♂, LEM); same data except, 13.vii.2011, CCBD-2141- D02 (1 ♂, barcoded, LEM); Masik Riv., Banks Is., 4–10.vii.1968, W. R.N. Mason (16 ♂, 11 ♀, CNC); Masik Riv., Banks Is., 14–31.vii.1968, G.E. Shewell (29 ♂, 19 ♀, CNC); Holman, Victoria Is., 26.vi.–10.vii.1952, D.P.Gray (3 ♂, 6 ♀, CNC); Hooper Is., Mackenzie Delta, 28.vi.–16.vii.1971, D.M. Wood (3 ♂, 5 ♀, CNC); Kidluit Bay, N. Richards Is. W., 27.vii.1948, J. R. Vockeroth (1 ♀, CNC); Sachs Harbour, 27–30.vi.1968, W. R. Mason (2 ♂, 2 ♀, CNC); Salmita Mines, 64°05′N, 111°15′W, 4.vii.1953, J.G. Chillcott (1 ♂, 4 ♀, CNC); - 21 mi E Tuktoyaktuk, 20.vi.–12.vii.1971, D.M. Wood (7 ♂, 2 ♀, CNC); Victoria Is., 71°17′N, 114°W, 1–28.vii.1975, G. & M. Wood (14 ♂, 11 ♀, CNC). Nunavut: Axel Heiberg Is., 79°25′N, 90°45′W, 19–31.vii.1963, H.K. Rutz (92 ♂, 54 ♀, CNC); Cambridge Bay, 12.viii.1950, W. R.M. Mason (4 ♂, 2 ♀, CNC); Cambridge Bay, 12.viii.1959, W. R. Richards (2 ♂, 2 ♀, CNC); Cambridge Bay, 18–31.vii.1950, E.H.N. Smith (2 ♂, 9 ♀, CNC); Cambridge Bay, 22–26.vii.1950, G.K. Sweatman (6 ♂, 9 ♀, CNC); Cambridge Bay, 69.120 7, -105.42582, 11–19.vii.2011, mesic, MT / sweep, Repl.2–3, NBP field party (4 ♂, LEM); Cambridge Bay, 69.121 57, -105.43124, 15–19.vii.2011, mesic, MT, Repl.3, NBP field party, CCDB-21421-G06, CCDB-21421-G09 (2 ♀, barcode associated, LEM); Chesterfield, 30.vii.–6.viii.1950, J. R. Vockeroth (1 ♂, 1 ♀, CNC); Chesterfield, 6–31.viii.1950, J.G. Chillcott (2 ♂, 1 ♀, CNC); Head of Clyde In- let, Baffin Is., 7.viii.1958, J.E.H. Martin (1 ♂, 5 ♀, CNC); Coppermine, 15.vii.1951, S.D. Hicks (2 ♂, 4 ♀, CNC); Coral Harbour, Southampton Is., 1–17.vii.1948, G. E. Shewell (55 ♂, 64 ♀, CNC); Coral Harbour, Southampton Is., 18.vii.1952, T.P. R. Ehlich (1 ♂, CNC); Eureka, 2–3.viii.1963, J. R. Vockeroth (4 ♂, CNC); Hot Weather Creek, Forsheim Peninsula, Ellesmere Is., 79°58′N, 84°28′W, 24.vi.–26.vii.1990, F. Brodo (13 ♂, 6 ♀, CNC); Eureka, 6–30.vii.1954, P.F. Bruggemann (3 ♂, 5 ♀, CNC); Fraser Lake, 68°45′N, 120°36′W, 19.vii.1969, G.E. Shewell (4 ♂, 5 ♀, CNC); Frobisher Bay, 2–8.vii.1948, F.G. DiLabio (2 ♀, CNC); Frobisher Bay, Baffin Is., 26.vi.1948, T.N. Freeman (1 ♂, CNC); Hazen Camp, 81°49′N, 71°18′W, 1–30.vii.1962, R.B. Madge (20 ♂, 19 ♀, CNC; 1 ♂, 1 ♀, UGIC); Hazen Camp, 81°49′N, 71°18′W, 1.vii.1962, P.S. Corbet (1 ♂, 1 ♀, CNC); Hazen Camp, 81°49′N, 71°18′W, 5–7.vii.1962, R.E. Leech (3 ♂, 3 ♀, CNC); Hazen Camp, 81°49′N, 71°18′W, 11–14.vii.1963, H.K. Rutz (2 ♂, 8 ♀, CNC); Hazen Camp, 81°49′N, 71°16′W, 12–26.vii.1962, J.A. Downes (8 ♂, 4 ♀, CNC); Hazen Camp, Ellesmere Is., 20.vii.–18.viii.1961, 14.vii.1962, D. R. Oliver (14 ♂, 12 ♀, CNC); Hazen Camp, 81°49′N, 71°16′W, 21.vi.1962, J.F. McAlpine (1 ♂, CNC); Hazen Camp, 81°49′N, 71°18′W, 1.viii.1963, J. R. Vockeroth (1 ♀, CNC); Hazen L., Ellesmere Is., 7–13.vii.1963, B. Hocking (6 ♀, CNC); Hazen L., Ellesmere Is., 13.vii.1963, C.D. Sharplin (3 ♀, CNC); Hazen L., Ellesmere Is., 27.vi.1967, J.D. Shorthouse (1 ♂, 1 ♀, CNC); Ellesmere Is., Lake Hazen, 81.82975°, -71.32244° [Repl.1], 81.82941°, -71.48788° [Repl.2], 19–25.vii.2010, wet & mesic, MT / sweep, NBP Field Party (7 ♂, LEM); Keewatin, 64°15′N, 89°30′W, 22.vii.1950, J.G. Chillcott (1 ♂, CNC); Lady Melville Lk., 93°15′W, 69°25′N, 3.vii.1951, J.G. Chillcott (1 ♂, 1 ♀, CNC); Melville Pen., Rose Bay, 66°51′N, 84°53′W, 19–22.vii.1948, Cody & McArthur (2 ♂, CNC); Kugluktuk, 67.83538°, -115.20987°, 3–7.vii.2010, mesic, MT, Repl.2, NBP Field Party (4 ♂, LEM); Nettilling Lake, Baffin Is., 7–9.vii.1925, J.D. Soper (2 ♀, CNC); Notting- ham Is., J. Mckenzie (1 ♂, USNM); Pond Inlet, Baffin Is., 26.vii., 6.viii.1978, 1.vii.1979, G. Johnston (3 ♂, 2 ♀, UGIC); Repulse Bay, 6.vii.1950, P.F. Bruggemann (1 ♂, CNC); Repulse Bay, 4–26.vii.1950, J.E.H. Martin (1 ♂, 4 ♀, CNC); Tanquary Fjord, Ellesmere Is., 18.vii.1963, H.K. Rutz (1 ♂, CNC); Head of Traquary Fjord, Ellesmere Is., 2.viii.1963, J. R. Vockeroth (2 ♀, CNC); Spence Bay, 2–30.vii.1951, J.G. Chillcott (12 ♂, 24 ♀, CNC); Spence Bay, 11–30.vii.1951, A.E. R. Downe (3 ♂, 13♀, CNC); Wager Bay, 65°15′N, 88°00′W, 22.vii.1950, J.G. Chillcott (1 ♂, 1 ♀, CNC). Yukon: British Mts, 69°27′N 140°25′W, 2–4.vii.1984, G. &M. Wood, D. Lafontaine (1 ♂, 1 ♀, CNC); Dempster Hwy, km 456, MT in tundra stream, 7–9.vii.1985, S.A. Marshall (3 ♂, UGIC); Firth River, 11– 14.vii.1956, R.E. Leech (8 ♂, 13 ♀, CNC); Firth River, 11–14.vii.1956, E.F. Cashman (3 ♂, 2 ♀, CNC); Herschel Is., 5–28.vii.1953, J.S. Waterhouse (24 ♂, 24 ♀, CNC); Herschel Is., 9–30.vii.1953, C.D. Bird (21 ♂, 29 ♀, CNC); Herschel Is., 28–29.vi.1971, D.M. Wood (12 ♂, 5 ♀, CNC); Herschel Is., 18–20.vii.1930, O. Bryant (1 ♂, 3 ♀, CNC); saddle, 68°01′N 136°42′W, 900 m, 1.vii.1987, J. Troubridge (1 ♂, 1 ♀, CNC). USA. Alaska: Cape Thompson, 22–26.vii.1961, R. Madge (14 ♂, 23 ♀, CNC); Kivalina R., 68°16′N 163°27′W, 10–17.vii.1974, K.W. Philip (2 ♂, CNC); mi. 213, Richardson Hwy, 17.vi.1951, J. R. McGillis (1 ♀, CNC); Schrader Lk., 3–5.vii.1972, K.W. Philip (1 ♂, CNC); St. George Is. [not shown on Fig. 16], 26–28.vii.1897, T. Kincaid (1 ♂, 3 ♀, USNM). GREENLAND. Auleitsivik, Nordskj. Exp. ’33 (1 ♂, USNM); Nedre Midsommer Sö [not shown on Fig. 16], 1–30.vii.1966, Can. Peary Land Expd. (86 ♂, 91 ♀, CNC); Sondrestrom Air Base, 10–14.vi., 18.vii.1952, W.J. Brown (10 ♂, 7 ♀, CNC); Sondrestrom Air Base, 11.vii.1963, W.P. DuBose (3 ♂, 3 ♀, USNM). Diagnosis. This species is distinguished from other Dasyrhamphomyia by the multi-serial acrostichals and dorsocentral setae; dark vittae beneath acrostichal and dorsocentral rows, separated by pair of lighter grey stripes; pale halteres; male cerci very long, pale, folded or wrapped over each other in dried specimens, leaving no opening dorsally and right cercus with mid-length expansion; male mid and hind tibiae and femora with pronounced ventral pile. Female wing darkly infuscate and broader than male. Redescription. Wing length 6.0–7.0 mm. Male. Head dark in ground-colour, with greyish pruinescence on face, frons, postgena and occiput; oral margin dark and lustrous. Holoptic, eye with ommatidia larger on upper half and smaller on lower half. Frons represented by very small triangular space below ocellar tubercle and larger subtriangular space above antennae, bare. Face divergent towards mouthparts, with oral margin dark and shiny. Ocellar triangle dark with 1 pair of anterior divergent setae, 1 pair of similar posterior setae; 2–3 pairs of postocellar setulae slightly shorter than ocellar setae. Occipital setae black, long, thin. Postgena with many long, hair-like dark setae. Antenna dark; scape slightly more than 2X longer than bulbous pedicel; pedicel broader than scape width; postpedicel cone-like, less than 3X basal width; stylus subequal to basal width of postpedicel. Palpus dark, with many dark, long, wavy hair-like setulae. Clypeus with upper margin glossy, grey pruinescent distally; labrum lustrous and reddish brown with yellow apex; labellum with dark setae. Thorax dark in ground-colour, largely densely grey-blue

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