Quedius meridiocarpathicus Smetana A. Solodovnikov 1958

Quedius meridiocarpathicus Smetana, 1958 (Figs 1, 2C, 4, 8B, 13F, 20) Quedius meridiocarpathicus Smetana, 1958: 365 [Type locality: Slovensko, Kamenný Most, Slanec] Quedius dalmatinus Korge, 1960b: 68 [Type locality: Dalmatia: Castelnuovo] References. Cඈංൿൿൺංඍ (1961): 53 (characters); KඈඋGൾ (1962): 153 (synonymy of Q. dalmatinus); Sආൾඍൺඇൺ (1962): 138 (characters); LඈHඌൾ (1964): 212 (characters); Hඈඋංඈඇ (1965): 277 (distribution); Sආൾඍൺඇൺ (1967): 558 (distribution); Bඈඋൽඈඇං (1976): 99 (characters); Cඈංൿൿൺංඍ (1978): 195 (characters); TඬඍH (1984): 129 (characters); Sආൾඍൺඇൺ (1993): 50 (distribution); Cංർൾඋඈඇං & Zൺඇൾඍඍං (1995): 32 (distribution); Lගඌඓඅඬ (1998): 137 (distribution); Aඇඅൺ෡ (2009): 230 (distribution); Sඈඅඈൽඈඏඇංĸඈඏ (2012): 33 (distribution); Pൾඍඋൾඇĸඈ (2015): 80 (distribution); Sൺඅඇංඍඌĸൺ & Sඈඅඈൽඈඏඇංĸඈඏ (2019): 49 (characters and distribution). Type material examined. Quedius meridiocarpathicus: Lൾർඍඈඍඒඉൾ J (cSme), designated by Sඈඅඈൽඈඏඇංĸඈඏ (2012), labelled: “ Slovakia mer. Kamen. Most 5.5.1955 Smetana 1955 / Quedius meridiocarpathicus s. Smetana det. 1957 / Lectotype Quedius meridiocarpathicus Smetana A. Solodovnikov des. 2009 / Quedius meridiocarpathicus Smetana A. Solodovnikov det. 2009”. Pൺඋൺඅൾർඍඈඍඒඉൾඌ: 1J, labelled:“ Slovakia mer. Kamen.most 3.5.1955 Smetana / Quedius meridiocarpathicus Smetana det. 1957 / Quedius meridiocarpathicus A. Solodovnikov det. 2009 / PARALECTOTYPE Quedius meridiocarpathicus SmetanaA.Solodovnikov des.2009 / NHMD 00523118” (NHMD); 1♀, labelled: “ Slovakia mer.Kamen.most 3.5.1955 Smetana / Quedius meridiocarpathicus Smetana det. 1957 / Quedius meridiocarpathicus A. Solodovnikov det. 2009 / PARALECTOTYPE Quedius meridiocarpathicus Smetana A.Solodovnikov des.2009 / NHMD 00523117”. Additional material examined. ALBANIA: Kolonjë, 3 km ENE Leskovik, 40.1586, 20.6216, flooded pasture, 990 m, 28.V.2010, M. Schülke (1 J cSch). BOSNIA AND HERZEGOVINA: Jajce, [44.34, 17.27], 12.-23.VI.1967, leg.Baderle (1 J 3♀♀ NMW); Trebinje, [42.71, 18.35], 1903 O. Leonhard (1 ♀ SDEI). BULGARIA: Burgas, Black Sea coast, [42.50, 27.48], 8.-19.V.1985, leg. D.W. Wrase (1 J 5 ♀♀ ZMHB); Kiten, [42.24, 27.77], 29.V.1982, leg. H. Wendt (1 ♀ ZMHB); Melnik, [41.52, 23.38], 12.-20.V.1981 (2 ♀♀ ZMHB); Nessebar, [42.66, 27.71], 6.-7.VI.1982, leg. H.Wendt (1 J 2♀♀ ZMHB); Pomorie, Black Sea coast, [42.58, 27.62], 9.-18.V.1985, leg. D.W. Wrase (1 J cSch); Primorsko, [42.26, 27.75], VII.1980, leg. I. Pokorný (1 ♀ cSch); Rupite, General Todorov, [41.43, 23.24], 5.V.1984, leg. Hieke (1 ♀ ZMHB); nr. Sandanski, [41.55, 23.23], 28.IV.1985, leg. B. Jaeger (1 J 1 ♀ ZMHB); Slivno, [42.67, 26.32] (1 J NHMD); Veliko Tarnovo, [43.08, 25.63], 25.VII.1984, leg. U. Opitz (2 ♀♀ ZMHB). CROATIA: Castelnuovo [Kaštel Novi], [43.55, 16.31], 1895, leg. F. Müller (1 ♀ SDEI); nr. Plitvice, [44.86, 15.58], 19.VII.1910, leg. Hochetlinger (1 ♀ NHMD); nr. Zagreb, [45.78, 15.98] (1 J NMW). GREECE: Alexandroupolis, [40.85, 25.87], 20.-30.IV.1967, leg. Krätschmer; 4 km NO of Anthili, Thessalia, Lamia, 38.8655, 22.5224, 6.II.2015, leg.T.Wolsch (1J cSch); Essymi [Aesymi], [41.01, 25.95], 700 m, 25.IV.1978, leg. Heinz (1 J 1 ♀ ZMHB); Ioannina, Metsovo, Lago O Pigon, [39.81, 21.11], 18.V.2005, 1500m, leg.Angelini (2 ♀♀ NHMD); Janina [Ioannina], [39.66, 20.85], IV.1927, leg.C. Purkyne (2 JJ NHMD); Keramidi, Thessalia, Palamas, 39.5627, 22.0801, 5.II.2015, leg. T. Wolsch (1 ♀ cSch); Lavrio, 60 km S Athina, [37.71, 24.06], 10.IV.1993, leg. P. Jørum (1 ♀ NHMD); Parnass. [Parnassos], [38.53, 22.62] (3 JJ NHMD); Skarfia, 20 km WSW Lamia, 38.8405, 22.6988, salt meadows, 6.II.2015, leg. T. Mainda (1 J cSch). Cඈඋൿඎ: Val di Ropa, [38.24, 21.73], 1905, leg. O. Leonhard (1 ♀ SDEI). Cඋൾඍൾ: Crete [35.41, 24.75], leg. Paganetti (1 J NMW) [potentially mislabelled or introduced specimen clearly belonging to this species]. Kൾൿൺඅඈඇංൺ: Paranetti, [38.21, 20.58], leg. F. Müller (1 ♀ SDEI). Pൾඅඈඉඈඇඇൾඌ: 3 km E Kalogria, 20 km NW of Varda, [38.19, 21.39], 15.IV.1993, leg. P. Jørum (1 ♀ NHMD); Nomos Lakonias, Near Fokiano, 37.0741, 22.9676, 0 m, sifting, waters edge, stagnant water, salt marsh, 02.IV.2019, leg. M. J. Justesen & A.K.Hansen (1 J NHMD). ZൺKඒඇඍHඈඌ: Kalamaki, [37.74, 20.89], 1909, leg. M. Hilf (1♀ ZMHB). HUNGARY: Hódmezővásárhely, [46.41, 20.34], 16.-22.VI.1995, leg Probst (2 ♀♀ NMW); Rapnitz [Rábca] river, Neusiedler See, [47.67, 16.81], V.1994 (1J ZMHB); Velencei, Dinnyes, [47.17, 18.54], salt lake with gravel banks, 15.VI.1979, leg. M. Uhlig (1 J 1♀ ZMHB). ITALY: Albareto, Emilia, [44.53, 10.40], 8.II.1994, leg.A. Fiori (2♀♀ ZMHB); NE of Bibione, left bank of Canale di Lugugnana mouth at Bibione, Veneto, 45.6259, 12.9791, sifting flood debris and litter, 29.V.2016, leg. A. Solodovnikov & M. Salnitska (2 JJ 1 ♀ NHMD); Casinalbo, Emilia, [44.53, 10.40] 25.VII.1996, leg.A. Fiori (2 JJ ZMHB); Cascine Del Riccio, Riva Ema [43.73, 11.25], 14.I.1970, leg. Mascagnia (1 J cBor); Comacchio, [44.69, 12.17], 28.III.1974, leg. P. Garaghahi (2 ♀♀ cBor); Firenzuola, [44.12, 11.37], VII.1970, leg. Mascagnia (1 J cBor); Lido di Jesolo, [45.51, 12.62], 10.VIII.1962, leg. Salvato (1 ♀ cBor); Noghera Istria, [45.58, 13.81] (1 J NHMD); Padova, [45.39, 11.90], IX.1969 (1 J ZMHB); Roma, Portonaccio, [41.90, 12.54], IX.1937, leg. G. Sacca (1 J 4 ♀♀ cBor); Tencarola, [45.39, 11.80], 10.XII.1961, leg. Salvato (1 ♀ cBor). NORTH MACEDONIA: Bitola, Jugoslavia, [41.02, 21.32], 1962, leg. J.T. Skovgaard (1 ♀ NHMD); Katlanovo nr. Skopje, [41.89, 21.68], 17.IX.-12.X.1966, leg. Gruber (1J 1♀ ZMHB); Treska River, Sara, [41.97 21.30], 3.VI.1980, leg.F. Hieke (1 J ZMHB); Vardar River, n. Skopje, [41.98, 21.49], 26.V.1980, leg. F. Hieke (1 J 3 ♀♀ ZMHB). ROMANIA: Predeal, [45.49, 25.57], 20.VI.1977 (1 ♀ MZLU). RUSSIA: Kඋൺඌඇඈൽൺඋ Kඋൺං: 15 km S of vill. Taman, [45.21, 36.70], 15.V.1995, sandy sea shore, under logs (2 JJ NHMD); Temruk, mud volcano Gefest, 45.2566, 37.4326, leaf litter in meadows, 12.VI.2017, leg. M. Salnitska (1 NHMD) Karabetova Gryada 5 km SE of vill. Taman, [45.12, 36.76], in litter at the bank of the permanent pond (1 ♀ NHMD); Mt.Tkhab, valley of river Zhene, [44.58, 38.35], in forest litter, 21.VI.1992, leg. M. Savitsky (1 J NHMD). SLOVENIA: 9 km SE Piran, [45.48, 13.59], 26.VI.1981, leg. F. Hieke (1 ♀ ZMHB). TURKEY: Adapazarı, [40.79, 30.40], flooding, 15.VII.1965, leg. Korge & Heinz (1 J ZMHB); SW Arnavutköy and Yassıören, [41.17, 28.65], 60 m, 19.IV.1990, leg. Heinz (1 J ZMHB); Caykasi River, Erfelek, Sinop, [41.89, 34.92], 26.V.1989, leg. Schönmann & Schillhammer (1 J 3 ♀♀ NMW); 6 km E Gebze, Izmit, [40.78, 29.49], IV.1983, leg. Heinz (1 J 2 ♀♀ ZMHB); 20 km W Gebze, [40.87, 29.22], IV.1989, leg. Heinz (2 JJ 1 ♀ ZMHB); Gelibolu, [40.41, 26.67], 10 m, 3.V.1982, leg. Boffa (2 ♀♀ cBor) İnönü, Eskişehir, [39.82, 30.14], 850 m, 3.IV.1986, leg. Heinz (1 ♀ ZMHB); Keşan, [40.85, 26.62], 22.IV.1978, leg. Heinz (2 JJ ZMHB) E Pamucak, Selcuk, [37.96, 27.27], swamp, 28.III.1986, leg. Heinz (1 J ZMHB); 2 km E Sarısaltık, 39.1241, 39.2475, 1594 m, 19.V.2011, leg. Anlaş (1 ♀ NHMD); Saray, 30 km W Ankara, [40.07, 32.61], 23.II.1973, leg. Exp. Nat. Mus. Praha (1 J NHMD); Söğüt, SE of Bilecik, [40.02, 30.18], 1200 m, 4.IV.1992, leg. Heinz (1 J 2 ♀♀ ZMHB); Susuz-Dag, Dokuz-Göl, Elmalı, [36.50, 29.76], 1700-2000 m, 20.VII.1971, leg. Heinz (1 J ZMHB). UKRAINE: Environs Odessa, right bank of Kujalnitskij estuary, [46.57, 30.73], 10.VI.2005, under stones, leg.A. Gontarenko (3 JJ 1 ♀ NHMD); Nikitsky botanical garden, [44.50, 34.23], by the sea, stones, 16.IX.1966 (1 J cRyv). Redescription. Measurements JJ (n = 7): HW = 1.73– 1.82 (1.78); HL = 1.47–1.56 (1.51); HL/HW 0.82–0.88 (0.85); PW = 2.27–2.47 (2.36); PL = 2.09–2.31 (2.19); PL/PW 0.89–0.95 (0.93); EW = 2.27–2.47 (2.36); EL = 2.09–2.24 (2.17); EL/EW 0.87–0.99 (0.92); EL/PL 0.92–1.05 (0.99); PW/HW 1.50–1.66 (1.56); forebody length 5.71–6.02 (5.86). ♀♀ (n = 3): HW = 1.64–1.76 (1.71); HL = 1.33–1.51 (1.43); HL/HW 0.81–0.86 (0.83); PW = 2.07–2.36 (2.21); PL = 1.96–2.13 (2.05); PL/PW 0.91–0.95 (0.93); EW = 2.16–2.38 (2.28); EL = 1.98–2.24 (2.12); EL/EW 0.90–0.97 (0.93); EL/PL 1.00–1.09 (1.03); PW/HW 1.52–1.56 (1.54); forebody length 5.27–5.78 (5.60). Medium sized, robust species; body light to dark brown (Fig. 8B). Head black, distinctly transverse, with eyes medium sized (EyL/TL = 1.50–1.67 (1.56)); microsculpture of transverse waves; no interocular punctures between anterior frontal punctures (cf. Fig. 6F); antennae pale reddish, antennomeres elongate; palpi pale reddish. Thorax: pronotum light to dark brown, slightly wider than long, wider than head, with microsculpture of transverse waves; three punctures in dorsal row and one to two in sublateral row with its posteriormost puncture reaching just beyond the level of first puncture of dorsal row; scutellum punctured and pubescent; elytra most often reddish brown rarely fully darkened, uniformly pubescent, slightly wider than long, roughly same length as pronotum; legs reddish brown with inner face of tibia darker. Abdomen light to dark brown, tergites uniformly punctured, without clear iridescence. Male. Aedeagus (Fig. 13F): paramere spatulate with clear proximal to medial attenuation and extending into paddle-like structure near apex, reaching just beyond apex of median lobe, with sensory peg setae forming a broad horseshoe shaped band of relatively large peg setae; median lobe broad with gentle constriction to a point at apex, on parameral side with two small teeth pointing slightly basad, positioned at level near basal level of peg setae band of paramere; internal sac without a continuation of C-sclerite. Differential diagnosis. Quedius meridiocarpathicus is very similar to other species in the molochinus -group with red elytra, viz Q. molochinus, Q. balticus, and Q. vicinus. It can be distinguished from Q. balticus by the pale base of the basal three antennomeres. Generally, it has paler middle antennomeres compared to Q. molochinus. Compared to Q. vicinus it is generally slightly darker, elytr

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