Lasiopogon puyallupi Mcknight & Cannings, 2020, stat. nov.

Lasiopogon puyallupi stat. nov. Lasiopogon willametti puyallupi Cole & Wilcox, 1938: 79. Diagnosis. (Fig. 33) A medium-sized dark species from sandy rivers and beaches of northwestern Washington and British Columbia; all setae black except white setae of beard, coxae, femur, and tergite sides; thorax tomentum ranges from deep brown to grey with strong dorsocentral stripes; tergites usually with basal brown bands. Epandrial halves moderately concave medially, with black setae; phallus dorsal carina with protruding apex swollen but slightly dorsoventrally flattened, pointing in an acute angle with the aedeagal tube. Ovipositor dark brown/black, acanthophorite spines black. Redescription. Body length ♂ 10.0– 11.1mm; ♀ 10.2–11.5mm. Head. HW ♂ 2.20–2.37mm; ♀ 2.16–2.48mm. FW ♂ 0.49–0.58mm; ♀ 0.52–0.57mm. VW ♂ 0.92–0.99mm; ♀ 0.93–1.05mm. VW/HW = ♂ 0.42–0.44; ♀ 0.41– 0.43. FW/VW = ♂ 0.51–0.59; ♀ 0.54–0.56. VD/VW = ♂ 0.12–0.16; ♀ 0.12–0.14. GH/GL = ♂ 0.40–0.45; ♀ 0.37–0.48. Face with silver-yellow tomentum; vertex with grey-brown tomentum, small bare patches between macroseta groups. Mystax, frontal, orbital, ocellar, and occipital setae black, occasionally a few ventral setae on mystax white; beard and labial setae white. Occipital macrosetae long and curved to right angle, longest (to 0.8mm) behind dorsolateral angle; lateral and ventral macrosetae shorter, straighter. Frontal and orbital setae fine and long; almost as long as ocellar macrosetae. Antennae. Black; setae black, no setae on postpedicel. Postpedicel long and rectangular; stylus broadly tapering. WPP/LPP = ♂ 0.26–0.31; ♀ 0.27–0.31. LAS/LPP ♂ 0.60–0.67; ♀ 0.54–0.59. Thorax. Prothorax brown/grey, setae white; postpronotal lobes grey, lateral angle ferruginous, setae brown. Scutum tomentum ranges from dark brown to light grey. Dorsocentral stripes dark brown, cuticle subshining; acrostichal stripes slightly darker than base tomentum. All setae and macrosetae of scutum black; notal setae longer than scape. Dorsocentral macrosetae fine, mixed with other setae, 7–11 anteriors (to 1.0mm), 3–5 posteriors. Postalars 2–3, with a few shorter setae; supra-alars 2–4; presuturals 3; posthumerals 1–2. Scutellar tomentum grey-brown; rim inflated, leaving a semi-circular line impressed inside the dorsal edge; apical macrosetae black, very abundant (10+ on each side) and mixed with many other long setae. Pleural tomentum grey-tan. Katatergite macrosetae black, 8–10, with a few weaker setae; katepisternal setae moderately long, white; anepisternal setae black, 6–9 relatively strong (to 0.8mm) and mixed with other short setae along posterior margin, a patch of short setae on dorsal margin of sclerite; anepimeron with 0–2 short wispy setae. Legs. Base color black except apex of femur ferruginous; tomentum grey. No coxal peg. Macrosetae black, except basal 60% of femur; finer setae white on femur, black on tibia and tarsi. Ventral setae on femur as long as or slightly longer than femur width; dorsolateral macrosetae mixed with finer setae, 8–12 on profemur difficult to distinguish from surrounding finer setae, more prominent on mesofemur (4–8) and metafemur (11–17). Protibia with ventral macrosetae 2–3 times longer than tibial width; ventromedial patch of dense short setae on protibia pale yellow; dark brown on metatibia. Claws reddish brown over basal 60%, apically black. Wings. Veins dark brown; membrane faintly brown from microtrichia, darker around crossveins at apex of cells br and bm. DCI = 0.33–0.43. Halter cream, with no spot. Abdomen. Male. Cuticle dark brown/black; brown tomentum at bases of tergites, bands of grey tomentum cover apical 40% of each tergite as a straight band, extending anteriorly along lateral margins. Dorsal setulae dark brown; lateral setae all white; lateral macrosetae on first tergite mostly black (4–7), 1–3 straw-colored. Sternite tomentum grey; setae straw-colored/white. Female. As in male, but grey tomentum extends anteriorly along midline to divide brown into lateral spots, and lateral setae on tergites 4–7 black; sternite setae sparser than in male. Male genitalia. Epandrium and hypandrium/gonocoxite complex black; sparsely covered in thin grey tomentum and black setae, setal brush dark brown, dense. Epandrium elongate, the width of the halves in lateral view about 50% the length, widest in basal third; apex rounded with ventral tooth. In dorsal view, medial margins of epandrium moderately curved, curvature starts about 30% from base, outer margins gradually tapered; basal sclerite prominent. Phallus paramere sheath dorsally about 48% the length of phallus; paramere sheath without ventral ornamentation; dorsal carina protruding, with apex swollen but slightly dorsoventrally flattened and pointing in an imaginary acute angle with the aedeagal tube. Ejaculatory apodeme in lateral view relatively straight, with broad dorsal carina along entire length. Subepandrial sclerite with triangular central unsclerotized area in basal 35% narrowing to a parallel-sided gap in central portion; spines parallel-sided, blunt, densely distributed basally, apically, and along midline, sparse laterally. Female genitalia. Undissected: Tergite 8 dark brown/black, with erect white setae; sternite 8 dark brown with short brown setae; lateral lobe setae strong, black; hypogynial valves with fine setae concentrated medially. Cerci brown/black with pale setae; acanthophorite spines black. Type Material. LECTOTYPE (here designated) ♂ labelled: “[rectangular white label] Auburn, Wn./ V-10 1932 / J. Wilcox, Col.”; “[rectangular beige label] PARATYPE / Lasiopogon / willametti/ Cole & Wilcox”; “[rectangular beige label] FORM/ puyallupi/ Cole & Wilcox”; “[rectangular white label] JOSEPH WILCOX/ COLLECTION – 1981/ Gift to California / Academy of Sciences ”. Our lectotype label “[red and black bordered label] LECTOTYPE / Lasiopogon puyallupi / Cole & Wilcox 1938 / des. T.A. McKnight 2017” has been added. Deposited at the CAS. PARALECTOTYPES (41 specimens designated): U.S.A.: WASHINGTON: King Co., Auburn [47.305 - 122.193], 10.v.1932, J. Wilcox (14♂ 1♀ CAS, 2♂ USNM, USNMENT1100230 – USNMENT1100231); Enumclaw [47.175 -122.029], 12.v.1932, J. Wilcox (10♂ CAS, 1♂ 1♀ [1 pr in cop] FISH); Pierce Co., Buckley [47.172 -122.039], 14.vi.1932, J. Wilcox (2♂ CAS), 15.vi.1932, J. Wilcox (1♂ CAS); Mt. Rainier, Ipsut Creek Camp [46.9755 -121.828], 26.vi.1932, W.W. Baker (4♂ 1♀ CAS); Sumner [47.203 -122.248], 10.v.1932, J. Wilcox (1♂ CAS), 17.vi.1932, J. Wilcox (1♂ CAS, 1♂ FISH, 1♀ USNM, USNMENT1100250). Other Material Examined (207 specimens). CANADA: BRITISH COLUMBIA: Agassiz [49.239 -121.767], 18.v.1927, H.H. Rose (1♀ CNC); Delta, Deas Island [49.125 -123.062], 26.iii.1983, R.A. Cannings (1♂ 1♀ RBCM, RBCM ENT 991-86355, RBCM ENT 991-86358), 6.iv.1984, R.A. Cannings (1♀ RBCM, RBCM ENT 991-86357), 10.iv.1991, R.A. Cannings & H. Nadel (1♀ RBCM, RBCM ENT 991-67087), 20.iv.1991, R.A. Cannings & H. Nadel (8♂ 1♀ RBCM, RBCM ENT 991-67067, RBCM ENT 991-67070, RBCM ENT 991-67079 – RBCM ENT 991-67081, RBCM ENT 991-67083, RBCM ENT 991-68073 – RBCM ENT 991-68074, RBCM ENT 991-68084), 12.v.1991, R.A. Cannings (1♂ 2♀ RBCM, RBCM ENT 991-67982 – RBCM ENT 991-67984), 7.v.1997, R.A. Cannings (1♂ CUAC, 6♂ 1♀ RBCM, 1♂ 1♀ TAM); Dewdney, River Road South [49.144 -122.182], 19.v.1991, R.A. Cannings (1♂ RBCM, RBCM ENT 991-68073); Fraser River, mouth of Hunter Creek [49.359 -121.575], 4.v.1989, R.A. Cannings (1♀ RBCM, RBCM ENT 991-1377); Hope, Coquihalla River, 49.383 -121.45, 13.vi.1995, R.A. Cannings (1♂ RBCM); Hope, Hunter Creek, at Highway #1 bridge, under RR bridge [49.358 -121.574], 23.v.1996, S.G. Cannings (1♂ RBCM); Ladner [49.092 -123.095], 23.iv.1987, S.G. Cannings (2♂ CSUC, 1♂ RBCM), 18.v.1991, R.A. Cannings (7♂ 1♀ RBCM, RBCM ENT 991-68051, RBCM ENT 991-68055, RBCM ENT 991-68060, RBCM ENT 991-68061, RBCM ENT 991-68063, RBCM ENT 991-68065, RBCM ENT 991-68066, RBCM ENT 991- 68068), 19.v.1991, R.A. Cannings (1♂ RBCM, RBCM ENT 991-68054); Lillooet River, Meager Creek Hotsprings, terrace [50.576 -123.465], 19.vii.1988, C.S. Guppy (1♂ RBCM, RBCM ENT 991-8089); Lillooet River, near John Sandy Creek [50.486 -122.944], 9.vi.1981, R.J. Cannings (1♂ RBCM, RBCM ENT 991-86356); Meager Creek road, 16.v.1987, C.S. Guppy (1♂ RBCM); Miracle Beach Provincial Park [49.85 -125.094], 28.v.1988, S.G. Cannings (1♂ LACM, LACM ENT 334064); Tofino, Chesterman Beach [49.114 -125.893], 27.v.1994, R.A. Cannings (2♂ CSUC, 1♂ LACM, LACM ENT 334063, 7♂ RBCM, RBCM ENT 010-002796, 1♂ TAM); Tofino, Wickaninnish Beach [49.013 -125.675], 16.v.1994, R.A. Cannings (1♂ 2♀ RBCM); Vancouver Island, Brooks Peninsula, Cape Cook Lagoon, 50.2 -127.8, 2.viii.1981, R.A. & S.G. Cannings (1♂ 1♀ RBCM, RBCM ENT 991-86364, RBCM ENT 991-86369), R.A. Cannings (1♂ 6♀ [1 pr in cop] RBCM, RBCM ENT 991-86366 – RBCM ENT 991- 86371); Vancouver Island, Comox, Cape Lazo Road, 49.68828 -124.87444, 2.v.2006, R.A. Cannings & R.G. Bennett (1♂ 1♀ RBCM). U.S.A.: OREGON: Hood River Co., E Fork Hood River at Nottingham campground, 6 mi E of Mt. Hood peak, 45.36815 -121.57066, 15.vi.2013, T.A.McKnight (1♀ [1 EtOH] UMMZ); WASHINGTON: Clallam Co., Bogachiel River and Sol Duc River confluence, Hwy 110 jct Mora Rd, 47.91311 -124.539, 23.vi.2010, R.A. Cannings (1♂ 1♀ RBCM, RBCM ENT 011-012688 – RBCM ENT 011-012689); Bogachiel River, Highway 101 [47.892 -124.355], 31.v.1995, R.A. Cannings & H. Nadel (2♂ RBCM, 1♂ 1♀ TAM); Forks [47.96 -124.393], 2.vii.1920, E.P. VanDuzee (2♂ 1♀ EMEC), 23.vii.1933, J. Wilcox (1♂ 2♀ CAS); Hoh River Campground, Hoh River, 47.85856 -123.93828, 24.vi.2010, R.A. Cannings (1♂ RBCM, RBCM ENT 011-012699), 25.vi.2010, R.A. Cannings (1♀ RBCM, RBCM ENT 011-012714); Quillayute River County Park, trail to Quillayute River bank, 47.91194 -124.58283, 23.vi.2010, R.A. Cannings (1♂ RBCM, RBCM ENT 011-011681); Cowitz Co., N Fork Toutle River, 10 mi E of Castle Rock, 1 mi S Hwy 504 jct Tower Rd, 46.3477 -122.7053, 14.v.2015, T.A. McKnight (1♂ RBCM); Grays Harbor Co.,

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