Tegoribates americanus Hammer. Setae (Roman 1958

Tegoribates americanus Hammer, 1958 (Figs. 9–16) Material examined: USA: Alaska, Fairbanks, 28.v.1948, (Lienk) 1 from sample 5 of Hammer (1955) (RNC); Denali N.P., Tolklat Ridge N.E., 29.viii.1976 (R. Greenberg) 4 from Carex and Allocomnium in willow area; Fort Yukon, 29.viii.1976 (S. Campbell) 12 from moss and alder litter; Atqasuk, Meade River IBP site, ‘Butterfly Creek’, 25.viii.1976 (A. Fjellberg) 8 from rich meadow high up on river bluff with Boykinia and Saxifraga; same location, 28.viii.1976 (A. Fjellberg) 5 from soil of S-facing slope with sandy rich dry meadow; California: San Louis Obispo Co., Cambria, 2.vii.1979 (DEW) 2 from Pinus radiata litter; Sacramento Valley, Jepson Prairie Reserve, 38°273N 121°825W, 13.iii.2009 (VBP, G. Pelletier) 4 from grassy area with Artemesia; Kansas, Riley Co., Konza Prairie, 3.xi.1981 (T. Seastedt) 1 from annually burned prairie soil; Canada: Yukon: Kluane N.P., 2500ft. elv., 28.vii.1983 (J. Robinson) 6; Richardson Mountains, Brown shale hill on Fish Creek, 68°N, 136°32’W, 2.vii.1987 (VBP) 1 from Hedysarum alpinum, Senecio, Saxifraga tricuspidata among rocks; Ogilvie Mountains, Km 155 Dempster Hwy, Windy Pass, 7.vii.1985 (VBP) 7 from Salix and Anemone at edge of stream; British Columbia, Cathedral Provincial Park, Glacier Lake, 2210m elv., 5.vii.1986 (VBP) 1 from mixed alpine vegetation, some lupins; same location, but 2286m elv., 7.vii.1986 (VBP) 10 from Draba ventosa, Sedum lanceolatum and Polemonium on shale above lake; Alberta: W Cynthia, 53.20847, -115.65059, 2010, (ABMI 2010.OG-1175-1SW) 1 adult; S Gypsy Lake Wildland, 56.31538, -110.5361, (ABMI 2007.544 NW) 1 adult; S Chinook Valley, 56.43682, -117.65735, (ABMI 2007.616 SE) 1 adult; SE Grand Rapids Wildland, 56.50687, -112.16037, (ABMI 2009.539 NW) 1 adult; W Millennium Lodge, 56.86831, -111.96440 (ABMI 2007.477 NE) 1 adult; N intersection Hwys 60 and 13, 52.50389, - 110.73015, (ABMI 2008.1224 NW) 1 adult; nr. Bicentennial Hwy., S Vermillion Lake, 59.86996, -112.57944, (ABMI 2009.13 SW) 1 adult; midway between Nipisi airport and Wabasca airport, 55.85966, -114.63938, (ABMI 2008.689 SE) 1 adult; W Bistocho Lake, 59.81076, -119.45875, 13.vi.2014 (ABMI 69 SW) 5 adults; S junction Hwys 264 and 322, 51.73390, -113.64149, (ABMI 2013 OG-1383 SW) 2 adults; between Hwys 50, 45, 670, 55.20903, -118.61084, (ABMI 2007.544 NW) 1 adult; (ABMI 2013.835 SW) 2 adults; ABMI 2016.63 SW, 28.v.2016, 2 adults; ABMI 2015.OG-583 SW, 28.v.2015, 7 adults; ABMI 2016.122 NW, 29.v.2016, 3 adults; NW Embarras airport, 17.vi.2013 (ABMI 238 NE) 2 adults; 57.24687 -112.51016, 27.v.2012 (ABMI 415 SE), 8 adults; W Conklin airport, 55.68718 -111.44270, 25.v.2012 (ABMI 668 SE) 6 adults. Revised diagnosis. Total length: 410–480. Longitudinal depressions ventrally extending from level of custodium to anal plate. Lamella about 104 long. Lamellar seta 17–20 long; mutual distance 20–26. Seta in about 20 long, smooth, acuminate; mutual distance 49–56. Bothridial seta fusiform, 90–94 long. Tutorium about 115 long, with distal tooth about 20 long, covering insertion of seta ro. Custodium tapered distally 3–5 long. Octotaxic system expressed as saccules, with Sa about 14 long, S1–S3 about 9 long. Description. Adult. Total length: 410–480; holotype, 410 (Hammer 1958); specimens from northern Canada considered conspecific by Hammer (1958), (n = 8) 440-460; total length based on specimens from California: females (n = 10) 469 (456–480); males (n = 3) 453 (448–456). Notogastral width females (n = 7) 309 (288–326); males (n = 1) 280. Integument. Integument smooth; with longitudinal depressions ventrally extending from level of custodium to anal plate (Fig. 10). Pteromorph, pedotectum I, gnathosoma, epimeres microtuberculate. Pedotectum I with fine striae along dorsal margin, pointed distally (Fig. 13A). Prodorsum. Seta ro about 41 long, strongly barbed along length, acuminate, curved anteromedially (Fig. 11). Lamella about 104 long, medially with V-shaped indentation; about 3 long. Lamellar seta arising anteriorly on fused lamella, just ventral of tip, 17–20 long; mutual distance 20–26. Seta in about 20 long, smooth, acuminate; mutual distance 49–56 (Fig. 9). Bothridial seta fusiform, 90–94 long (Fig. 9). Seta ex about 4, (Fig. 11; often not evident, but alveolus present). Dorsophragmata fused at base, about 34 long (Fig. 9, arrow). Lateral region of podosoma. Tutorium narrow tectum, about 115 long, with tooth distally, about 20 long, covering insertion of seta ro (Figs. 11, 13A). Custodium tapered distally 3–5 long (Fig. 10). Notogaster. Notogaster slightly longer than wide: 1.2:1; anterior margin straight. Lenticulus present. Octotaxic system expressed as saccules (Fig. 9), with Sa about 14 long, S1–S3 about 9 long, length difficult to measure, variable depending on orientation. Notogastral setae smooth, 15–23 long, setae subequal in length; lm usually medial and slightly posterior to Sa (Fig. 9); lp medial to S1, dorsocentrally positioned. Ventral. Epimeral setae mostly about 7–18 long, thin, smooth, acuminate, with 1c slightly barbed. Genital, aggenital, anal and adanal setae smooth; genital setae about 20, others about 11. Lyrifissure iad at anterolateral margin of anal plate. Postanal porose area about 36 long and 5 wide. Gnathosoma. Small axillary saccule present at base of palp, 5 x 2 (Fig. 13C). Length of seta m>a>h. Legs. Leg setal formulas I–IV, solenidia in parentheses: I: 1-5-3(1)-4(2)-20(2); II: 1-5-3(1)-4(1)-15(2); III: 2- 3-1(1)-3(1)-15; IV: 1-2-2-3(1)-12. Setae l” of genua and tibiae I and II thicker in shape than other setae on these segments, those of genua I and II strongly spinous (Figs. 12A, B). Seta pv” of tarsus II thick, strongly barbed (Fig. 13D). Tridactylous. Description Immatures dimensions: Mean length (based on slide-mounted specimens): larva (n = 3) 265 (239– 288); protonymph (n = 1) 305; deutonymph (n = 2) 360, 436; tritonymph (n = 4) 473 (456–506). Length of prodorsal and hysterosomal setae given in Table 1. Larva (Fig. 14). Integument unsclerotized; tuberculate; that of hysterosomal venter more strongly tuberculate than dorsum. Hysterosomal setae borne on tubercles. Prodorsum: Prodorsal setae, other than bothridial seta, all short, smooth, tapered; mutual distance of setae roro, le-le and in-in about 15, 24 and 35, respectively. Bothridial seta setiform, tapered distally. Gastronotic Region: Tubercle posteriorly bearing setae dp largest of hysterosomal tubercles. Gastronotal setae of c, d, series and la, lm, h3 and h1, short, smooth, tapered; lp barbed, brush-like, blunt distally; h2, long, barbed, almost spinous. Mutual distance of setae da-da, dm-dm and dp-dp almost subequal, about 11–13. Ventral Region: Medial margin of epimeral plates well defined. Epimere I with broad mentotectum, with medial edges well separated medially. Epimeral setae slightly barbed, isodiametric, about 8 long, except for scaliform seta 1c. Porose integument surrounding opening of opisthonotal gland not evident. Protonymph (Fig. 16C). Integument: Unsclerotized with light brown coloration; bearing larval scalp in 3 of 4 specimens examined. Venter of hysterosoma strongly tuberculate; centrodorsal region of hysterosoma almost smooth (as for tritonymph Fig. 15). Setae of l and h series borne on large tubercles; seta dp borne on medial tubercle. Prodorsum: Prodorsal setae, other than bothridial seta, all short, smooth, tapered; mutual distance of setae roro, le-le and in-in about 18, 18 and 40, respectively. Bothridial seta setiform, tapered distally. Gastronotic Region: Tubercle posteriorly bearing setae dp (Fig. 16B). Gastronotal setae of c, d, series and la, lm, h1, h2, short, smooth, tapered; lp barbed, brush-like, blunt to rounded distally; h3 similar in shape to lp. Mutual distance of setae da-da, dm-dm and dp-dp, about 20, 23 and 10, respectively. Ventral Region: Medial margin of epimeral plates well defined. Epimere I with broad mentotectum well separated medially. Epimeral, genital, aggenital and adanal setae very slightly barbed, isodiametric, about 8 long. Porose integument surrounding opening of opisthonotal gland not evident. Gnathosoma: Axillary saccule present. Palpal eupathidium acm attached to solenidion along distal threequarters of solenidial length. Deutonymph (Figs. 16A, B). Integument: Unsclerotized with light brown coloration; bearing larval and protonymphal scalps in 1 of 2 specimens examined. Hysterosoma strongly tuberculate, except for region of hysterosoma surrounding setae dm, dp almost smooth (as for tritonymph Fig. 15). Setae of l and h series borne on large tubercles; seta dp borne on medial tubercle. As for protonymph, except for overall size. Setae ro le, in and ex tapered, smooth; Mutual distance of ro-ro, lele and in-in about 23, 24, and 54, respectively. Bothridial seta setiform, tapered. Gastronotal setae of c, d and p series and h1 smooth, tapered; other gastronotal setae brush-like, expanded distally. Centrodorsal region of hysterosoma shorter than in protonymph or larva. Tritonymph. Integument: Unsclerotized with light brown coloration; bearing larval, protonymphal and deutonymphal scalps in 4 specimens examined. Hysterosoma strongly tuberculate, except for region of hysterosoma bearing d setae almost smooth (Fig. 15). Setae of l and h series borne on large tubercles; seta dp borne on medial tubercle. As for deutonymph, except for overall size. Mutual distance of ro-ro, le-le and in-in 24–30, about 31, and 49– 59, respectively. Bothridial seta setiform, tapered. Gastronotal setae of c, d and p series short, smooth, tapered; other gastronotal setae brush-like, expanded distally, and tapered. Centrodorsal region of hysterosoma shorter than in deutonymph, protonymph or larva (compare figs. 14, 15, 16B). Scalps forming triangular cap posteriorly on hysterosoma, with larval setae of d series closely adjacent at pinnacle of cap; triangular cap surrounded by cluster of brush-like l and h series setae (Fig. 16D). Gnathosoma:

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