Lasioglossum (Dialictus) angelicum Gardner & Gibbs 2023, sp. nov.

Lasioglossum (Dialictus) angelicum sp. nov. urn:lsid:zoobank.org:act: 7D4DB588-1714-46C7-AAC0-8E44E2AF44A5 Figs 3–5, 76A, 78C, 86B, 87A, 90B, 92B, 93B, 110B Diagnosis Females of L. angelicum sp. nov. have the tegula very large (exceeding posterior margin of mesoscutum in dorsal view) and densely punctate (IS <1 PD), with inner posterior margin strongly concave and forming a posterior projection about the size of 1 lateral OD; scutellum densely punctate (IS <1 PD); metapostnotum shiny with strong anastomosing rugae reaching posterior margin; T1 shiny and distinctly, moderately densely punctate (IS = 1–2 PD), usually with some fine punctures extending onto apical rim; face short (length/width ratio ≤0.83); gena slightly shorter than eye in lateral view; and tegula and metasoma black. Often, the mesoscutum is relatively sparsely and coarsely punctate (posterior half with IS = 1–3 PD in centre and only 2–3 punctures present between posterior end of parapsidal line and lateral edge of mesoscutum) and shiny at least on lateral and posterior margins, but a few specimens have the mesoscutum dull and more densely punctate; these specimens are difficult to distinguish from L. helianthi. Females of L. angelicum sp. nov. are most similar to those of L. gaudiale, L. helianthi, L. pseudotegulare, and L. tegulariforme. Females of L. gaudiale and L. helianthi have the tegula slightly smaller (reaching but not exceeding posterior margin of mesoscutum in dorsal view) and sometimes more sparsely punctate (IS ≥ 1 PD laterally at midlength). In addition, females of L. gaudiale have the gena about as wide as eye in lateral view, scutellum often moderately sparsely punctate (IS = 1–2 PD), and metapostnotum usually with subparallel rugae not reaching posterior margin. Females of L. helianthi have the T1 anterior slope usually coriarious and mesoscutum posterior half entirely densely punctate (IS <1 PD). Females of L. pseudotegulare have the face slightly longer (length/width ratio ≥ 0.83) and metapostnotum dull with fine, subparallel rugae. Females of L. tegulariforme have T1 with an impunctate median line about 1 OD wide and mesoscutum posterior half entirely densely punctate (IS <1 PD). Some females of other, unknown species in the L. stictaspis species complex can be very similar, but all have the T1 disc more finely and sparsely punctate (IS = 1–4 PD) and/or coriarious, and often have the tegula and metasoma red-orange. Males of L. angelicum sp. nov. have the tegula very large (as in the female); metapostnotum shiny with strong rugae; mesepisternum densely punctate (IS <1 PD) at least in ventral half; mesoscutum usually moderately densely punctate (IS = 1–2 PD); T1 shiny; discs of T1–T2 uniformly finely and moderately densely punctate (IS = 1–2 PD) with some punctures extending onto apical rims; face short (length/width ratio ≤ 0.86); and tegula black. Males of L. angelicum sp. nov. are most similar to those of L. helianthi, L. pseudotegulare, and L. tegulariforme. Males of L. helianthi have the tegula smaller (not exceeding posterior margin of mesoscutum in dorsal view) with a narrow posterior point much smaller than 1 lateral OD and mesoscutum densely punctate (IS ≤ 1 PD). Males of L. pseudotegulare have the face slightly longer (length/width ratio usually>0.86) and discs of T1–T2 more densely punctate (IS ≤ 1 PD). Males of L. tegulariforme have the disc of T1 more sparsely punctate than T2, T1–T2 apical rims impunctate, and mesoscutum densely punctate (IS ≤ 1 PD). Some males of other, unknown species in the L. stictaspis species complex can be very similar, but usually have the mesepisternum ventral half more sparsely punctate (IS = 1–2 PD), T1 coriarious at base, or tegula red-orange. A few specimens may be morphologically indistinguishable and can only be separated by geography, DNA barcodes, or associated females. Etymology The specific epithet ʻ angelicum ʼ is a Latin adjective derived from the noun ʻ angelus ʼ (angel) plus the adjectival suffix ʻ -icum ʼ (having the quality of). It is a reference to the type locality of the Los Angeles basin, where this species is common. An appropriate translation would be the angelic sweat bee. Material examined Holotype UNITED STATES – California • ♀; Los Angeles Co., Expo Park Nature Gardens, NHMLA; 34.02° N, 118.29° W; elev. 54 m; 18–20 Jul. 2018; H. Hayes leg.; LACM ENT 376858. [Verbatim label: USA: CA, Los Angeles / Expo Park Nature Gardens, / NHMLA 34.02°N 118.29°W / 54m, 18–20 July 2018, / H. Hayes 408 // LACM ENT 376858 // BOLD / gard0266- CA / DLIII 216-20 // HOLOTYPE / Lasioglossum (Dialictus) angelicum Gardner and Gibbs] Paratypes UNITED STATES – California • 1 ♀; Los Angeles Co., 5700 Tobias Avenue; 34.1739° N, 118.4518° W; elev. 209 m; 3 May 2018; S.B. Lerman leg.; ASUHIC LA-WF-2-BB-2 881 • 1 ♀; Los Angeles Co., 5831 Norwich Avenue; 34.1767° N, 118.4611° W; elev. 212 m; 11 Apr. 2018; S.B. Lerman leg.; ASUHIC LA-XS-3-BB-1 496 • 1 ♀; Los Angeles Co., Atwater Village; 34.114° N, 118.251° W; 28 May–4 Jun. 2014; Sidney Higgins leg.; LACM ENT 394539 • 1 ♂; Los Angeles Co., Ballona Wetlands near Playa del Rey; [33.97° N, 118.44° W]; 16 Jun. 1981; R.R. Snelling leg.; CUIC • 4 ♀♀, 1 ♂; Los Angeles Co., Chatsworth; 34.26° N, 118.601° W; 1–8 May 2017; LACM • 21 ♀♀, 5 ♂♂; ibid.; 1–8 Jul. 2017; LACM • 3 ♀♀, 5 ♂♂; Los Angeles Co., Claremont; 34.0894° N, 117.7111° W; 1–8 Jul. 2016; LACM • 4 ♀♀, 3 ♂♂; ibid.; 1–8 Aug. 2016; LACM • 1 ♀; Los Angeles Co., Eagle Rock; 34.129° N, 118.215° W; 3–10 May 2014; Eric Keller leg.; LACM ENT 394537 • 1 ♀; Los Angeles Co., Eagle Rock; 34.13° N, 118.23° W; elev. 169.9 m; 15–21 Jul. 2018; Hannah Hayes leg.; LACM ENT 380968 • 1 ♀; Los Angeles Co., Expo Park; 34.018° N, 118.289° W; elev. 55 m; 30 May–6 Jul. 2014; D. Pentcheff, L. Gonzalez leg.; LACM ENT 394536 • 2 ♂♂; same locality as for holotype; 18 Jul. 2018; Hannah Hayes leg.; LACM ENT 376863, 376871 • 1 ♀; same locality as for holotype; 13–15 Aug. 2018; Hannah Hayes leg.; LACM ENT 381174 • 14 ♀♀, 3 ♂♂; Los Angeles Co., Gardena; 33.876° N, 118.288° W; 2–9 Oct. 2018; LACM • 5 ♀♀, 1 ♂; ibid.; 1–8 Nov. 2018; LACM • 9 ♀♀, 1 ♂; Los Angeles Co., Gardena; 33.8843° N, 118.2866° W; 1–8 Jul. 2016; LACM • 1 ♂; Los Angeles Co., Glendale; 34.159° N, 118.247° W; 31 May–7 Jun. 2014; Celeste Armstrong leg.; LACM ENT 394543 • 1 ♀; Los Angeles Co., Mid-City; 34.047° N, 118.334° W; 31 May–7 Jun. 2014; Sharon Oxborough leg.; LACM ENT 394538 • 2 ♂♂; Los Angeles Co., Pasadena; [34.15° N, 118.14° W]; 24 Jun. 2000; John S. Ascher leg.; AMNH • 2 ♀♀; Los Angeles Co., Pasadena, Arroyo Park; [34.15° N, 118.165° W]; 7 Mar. 2001; John S. Ascher leg.; AMNH • 1 ♀; Los Angeles Co., Silverlake; 34.093° N, 118.274° W; 31 May–7 Jun. 2014; Walter Renwick leg.; LACM ENT 394540 • 11 ♀♀; Los Angeles Co., Torrance; 33.8712° N, 118.331° W; 13–23 Jul. 2017; LACM • 6 ♂♂; ibid.; 13–23 Jul. 2017; LACM • 1 ♂; Los Angeles Co., University Park; 34.034° N, 118.281° W; 28 May–4 Jun. 2014; Peggy Hentschke leg.; LACM ENT 394542 • 22 ♀♀, 11 ♂♂; Los Angeles Co., Valley Village; 34.171° N, 118.398° W; 1–8 Jul. 2017; LACM • 3 ♀♀, 1 ♂; Riverside Co., Norco; 33.912° N, 117.5702° W; 1–8 Jun. 2016; LACM • 1 ♀; San Diego Co., 2 km E of Pine Valley; 32.8247° N, 116.5042° W; elev. 1300 m; 31 May–5 Jun. 2002; F.D. Parker, M.E. Irwin leg.; BBSL FDP195086 • 1 ♀; Santa Barbara Co., CCBER Greenhouse, 61M NW Parking Lot 38, UCSB; 34.4209° N, 119.8581° W; 14 May 2019; CCBER UCSB-IZC00036859 • 1 ♀; Santa Barbara Co., Isla Vista, San Joaquin Restoration Project; 34.4177° N, 119.8655° W; 8–10 Mar. 2016; Elaine Tan leg.; CCBER UCSB-IZC00034561 • 1 ♀; Santa Barbara Co., UCSB North Parcel; 34.4224° N, 119.8794° W; 7 Feb. 2019; CCBER UCSB-IZC00036899 • 1 ♂; Ventura Co., Santa Clara River; 34.3891° N, 118.8853° W; 20 May 2019; CCBER UCSB-IZC00037014 • 1 ♀; Newport Beach; [33.62° N, 117.93° W]; 19 Aug. 1920; Philip H. Timberlake leg.; ex anise; UCRC ENT 525678 • 1 ♂; Riverside; [33.97° N, 117.32° W]; 31 Aug. 1924; Philip H. Timberlake leg.; ex on wing; UCMC • 1 ♀; ibid.; 10 Feb. 1925; Philip H. Timberlake leg.; ex Encelia farinosa; UCMC • 1 ♀; San Diego; [32.71° N, 117.16° W]; 22 Apr. 2004; DB leg.; WRME 518949. Description Female COLOURATION. Head and mesosoma blue to blue-green; clypeus apex black; labrum black; mandible orange with black base and red apex; flagellum black dorsally, orange ventrally. Pronotal lobe black; legs black with femur-tibia joints and tarsi dark reddish brown; tegula dark reddish brown; wing membrane hyaline, veins with subcosta dark brown, otherwise lighter brown. Metasoma black with rims of terga and sterna broadly translucent yellow. PUBESCENCE. Body pile colour white to pale yellow. Tomentum dense on lower paraocular area, gena adjacent to compound eye, pronotal collar and lobe, space between pronotal lobe and tegula, T2–T3 basolaterally, and T4 throughout. Mesoscutum pubescence thin to moderately plumose. Wing setae dark, short and dense. Acarinarial fan complete, dense. T2 fringes dense, T3 fringes dense. SURFACE SCULPTURE. Clypeus shiny, with punctures moderately sparse (IS = 1–2 PD), becoming larger in apical half; supraclypeal area shiny, with punctures moderately dense (IS = 1–2 PD); paraocular area shiny, with punctures dense (IS <1 PD), sparser around antenna socket (IS ≤ 1 PD); frons reticulate, with punctures fine, crowded (IS = 0 PD); vertex shiny, with punctures dense laterally (IS <1 PD), sparse medially (IS = 1–3 PD); gena shiny, with punctures fine, sparse (IS = 1–3 PD); postgena lineolate, becoming shiny anteriorly. Tegula punctures dense (IS <1 PD); mesoscutum weakly tessellate, with punctures moderately dense (IS = 1–2 PD), becoming dense on lateral and posterior margins (IS <1 PD); scutellum weakly tessellate, with punctures dense marginally (IS <1 PD), moderately sparse submedially (IS = 1–2 PD); metapostnotum shiny, becoming tessellate posteriorly, with rugae strong, anastomosing, reaching posterior margin; preëpisternum rugulose;

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