Sclerocoelus chilensis Kuwahara, Marshall & Paiero, 2025, sp. nov.
Bibliographic record
Abstract
Sclerocoelus chilensis sp. nov. urn:lsid:zoobank.org:act: 12116740-67B2-4A52-BCB6-A6E1F4C79981 Figs 5I, 32–33 Etymology This name reflects the apparent restriction of this species to Chile, where it is the only known species of Sclerocoelus. Material examined Holotype CHILE • ♂; Valparaíso, Portillo, Río Juncalillo; 32°50′58″ S, 70°07′59″ W; 2500 m a.s.l.; 10 Dec. 2008; Kits and Marshall leg.; stream vegetation; MNNC debu00310625. Paratype CHILE – Valparaíso • 1 ♂; same data as for holotype; DEBU debu00310638. Description BODY (Fig. 32A). Length 3.0– 3.1 mm. Head dark brown, ventrolateral corners of frons orange; gena slightly reddish. Frontal width 2.5× frontal height. Three pairs of strong interfrontal bristles surmounting a fine fourth pair; anterior orbital 0.4–0.5× length of posterior. Palpus yellow. Eye reduced, greatest height about 2.0× shortest genal height. Thorax dark brown, scutum with slightly paler lateral edges. Two pairs of dorsocentral bristles (anterior pair distinct, 0.5× length of posterior pair) separated by 7–8 rows of acrostichal setulae. Membrane around prosternum darkened but bare. Legs brown, femora apically and tibiae basally paler. Fore femur with 3–4 large ventral preapical setae. Dorsal surface of mid tibia lacking small anterior seta in distal half. Ventral surface of male mid tibia with two rows of stout setae along apical half. Wing (Fig. 5I) slightly infuscate. CS2 subequal to CS3. Halter brown. MALE ABDOMEN (Figs32B–C, 33).Dark brown, posterior edges of tergites sometimes slightly desclerotized. T2–5 and S2–4 uniformly long-setose with large posterolateral setae. S5 broad, asymmetrical, medially 1.0× length of S4, left side reduced anteriorly and posteriorly, setose in posterior half with a short, darkened, setulose posteromedial patch. Anterior flange of S6+7 not developed. Sclerite A large, rectangular, setulose; sclerite B very large and flat, curved to right side and laying parallel to S5; sclerite C triangular, fused to right extension of S6+7, situated over and to the left of sclerite D+E; sclerites D and E fused, triangular, posteriorly split with a slight overlap of apices; sclerite F large, setulose with a dark, sinuate apodeme on left side; sclerite G large, bulging, broadly fused to right side of sclerite F; ring sclerite well developed. Epandrium large, 0.8× length of S8, height 1.4 × maximum length and 0.7× maximum width, uniformly long-setose; perianal pads weakly developed. Pseudocercus absent; halves of subepandrial sclerite broad, strongly arched, and medially fused. Subcercus large, basal half broad in posterior view and angled anteroventrally in lateral view, apical half flattened in posterior view, tapered and angled posteroventrally in lateral view. Hypandrium with thin, sinuate anteromedial apodeme. Surstylus large, triangular, long-setose with a blade-like, inner anterobasal lobe and a curved, preapical, posterodorsal emargination. Postgonite large, strongly sinuate, and evenly tapered. Phallapodeme very large with a short dorsal ‘fin’; basiphallus stout and connected to distiphallus by a neck-like distal part; distiphallus short, largely membranous with a broad, sinuate dorsal sclerite and a pair of sinuate lateral sclerites. FEMALE ABDOMEN. Female unknown. Distribution Neotropical: Chile. Remarks Sclerocoelus chilensis sp. nov. is one of two species that has a darkened membrane around the prosternum, the other being S. rectangularis. Sclerocoelus chilensis can be readily separated from S. rectangularis by its smaller eye, smaller epandrium with an ovoid anal fissure, subtriangular subcercus, and bilobed genital pouch sclerite D. Sclerocoelus chilensis, the southernmost species of the genus, is known from small deposits of plant material along a stream in an otherwise dry and barren alpine landscape. An apparent restriction to the moist microhabitat created by bits of vegetation along high Andean streams or similar high elevation microhabitats is a characteristic of several related species, including S. lutosus sp. nov., S. punensis sp. nov., and S. tridens sp. nov.
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How this classification was reachedexpand
Full frame distilled prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. Learned from the 10,348 direct Codex labels and 10,348 direct Gemma labels. Candidate is the union of thresholded teacher heads; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels or direct frontier model labels.
Codex and Gemma teacher scores by category
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.001 | 0.000 |
| Meta-epidemiology (narrow) | 0.000 | 0.000 |
| Meta-epidemiology (broad) | 0.000 | 0.000 |
| Bibliometrics | 0.000 | 0.001 |
| Science and technology studies | 0.002 | 0.000 |
| Scholarly communication | 0.001 | 0.000 |
| Open science | 0.001 | 0.000 |
| Research integrity | 0.000 | 0.000 |
| Insufficient payload (model declined to judge) | 0.043 | 0.011 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from itClassification
machine, unvalidatedMachine predicted; both teacher heads agree on what is shown here.
How this classification was reached, model by model and score by score, is at the end of the page under "How this classification was reached".