Rudolfina exuberata Paiero & Marshall 2020, sp. nov.

Rudolfina exuberata sp. nov. urn:lsid:zoobank.org:act: C4509588-8528-4024-A87C-67A845A8157E Figs 1B, 9–10, 21D Etymology The specific name, which has been a manuscript name since Marshall (1982), is from the Latin for ‘abundant’: of the 1933 specimens examined for this study, 1649 were R. exuberata sp. nov. Material examined Holotype UNITED STATES OF AMERICA • ♂; Florida, Marion Co., Ocala National Forest; 14–18 Jun. 1984; S.A. Marshall leg.; creekside, dung; DEBU debu01087005. Paratypes ARGENTINA • 4 ♂♂; Misiones, 5 km E of Puerto Iguaz; 2–7 Feb. 1992; S.A. Marshall leg.; FIT/pans/ dung pans; DEBU. ú BELIZE – Cayo • 4 ♂♂, 2 ♀♀; Caves Branch; 23–29 Aug. 1972; S. and J. Peck leg.: forest, dung; DEBU • 2 ♂♂; Belmopan; 26–30 Jul. 1972; S. and J. Peck leg.; palm forest, carrion; DEBU • 2 ♂♂, 3 ♀♀; San Ignacio, Maya Mountain Lodge; 17°09′ N, 89°04′ W; 2 Jan. 1991; S.A. Marshall leg.; Atta mound trail; DEBU • 1 ♂, 2 ♀♀; San Ignacio, Maya Mountain Lodge; 17°09′ N, 89°04′ W; 7–8 Jan. 1991; S.A. Marshall leg.; dung trap; DEBU • 1 ♂; San Ignacio, Maya Mountain Lodge; 17°9′ N, 89°4′ W; 7 Jan. 1991; S.A. Marshall leg.; Malaise; DEBU • 1 ♂, 11 ♀♀; Mountain Pine Ridge, Hidden Valley Inst.; 2500 ft a.s.l.; 10–15 Jan. 1991; S.A. Marshall leg.; broadleaf forest, dung traps; DEBU • 1 ♀; Mountain Pine Ridge, Hidden Valley Inst.; 2500 ft a.s.l.; 14 Jan. 1991; S.A. Marshall leg.; pine–jungle edge, sweep; DEBU • 1 ♂, 1 ♀; Mountain Pine Ridge, Hidden Valley Inst.; 2500 ft a.s.l.; 14 Jan. 1991; S.A. Marshall leg.; dung traps in pine; DEBU • 10 ♂♂, 18 ♀♀; Mountain Pine Ridge, Hidden Valley Inst.; 2500 ft a.s.l.; 10–15 Jan. 1991; S.A. Marshall leg.; grass–pine dung traps; DEBU • 3 ♂♂, 1 ♀; Beaver Dam, 39 mi. W of Belize City; 6–10 Aug. 1972; S. and J. Peck; swamp forest, dung trap; CNCI. BOLIVIA – La Paz • 1 ♀; Heath River Wildlife Centre, ~ 21 km SSW of Puerto Heath; 12°40′ S, 68°42′ W; 5–7 May 2007; Marshall and Kits leg.; treefall, yellow pans; DEBU • 2 ♂♂, 1 ♀; Heath River Wildlife Research Centre; 12°40′ S, 68°42′ W; 5–9 May 2007; Paiero and Kits leg.; treefall, yellow pans; UASC • 1 ♀; San Antonio, ca 8 km S of Mapiri; 15°20′56″ S, 68°13′31″ W; 11 Apr. 2001; S.A. Marshall; secondary forest, dung pans; DEBU. BRAZIL – Minas Gerais • 1 ♂, 1 ♀; 1 km E of Lavras; 18–20 Feb. 1990; S.A. Marshall leg.; dung traps in ditch; DEBU. – Paraná • 4 ♂♂, 2 ♀♀; Londrina; 1–2 Feb. 1990; S.A. Marshall leg.; carrion pan traps; MZSP • 2 ♂♂, 4 ♀♀; Londrina, Mata dos Godoy; 28–31 Jan. 1990; S.A. Marshall leg.; MZSP • 1 ♂, 1 ♀; 30 km SE of Curitiba, BR 277; 6–9 Feb. 1990; S.A. Marshall leg.; dung traps; DEBU. – Rio de Janiero • 3 ♂♂, 1 ♀; Tijuca Forest Reserve; 1–28 Feb. 1990; S.A. Marshall leg.; Malaise; MZSP • 1 ♂; Nova Friburgo, Sítio Edelweiss; 26 Jan. 1990; S.A. Marshall leg.; Malaise; DEBU. – São Paulo • 1 ♂; Estación Biologica Boracea; 2 Dec. 2008; G.F.G. Miranda leg.; dung trap; MZSP • 1 ♀; Sao Paulo, Jaragua; 8 Feb. 1990; S.A. Marshall leg.; DEBU • 8 ♂♂, 7 ♀♀; USP Biology Station; 5–6 Feb. 1979; R. Woodruff and J. Runnacles; human dung; MZSP. CHILE • 2 ♀♀; Valparaíso, La Campana National Park; 22 Nov. 2006; S.A. Marshall leg.; MNNC. COLOMBIA • 2 ♂♂, 2 ♀♀; N of Santander, 20 mi. S of Cucuta Quebrada de Honda; 2500 ft a.s.l.; 13–15 May 1974; S. Peck leg.; carrion trap; DEBU • 1 ♂, 1 ♀; 2 mi. N of Chinacota; 3000 ft a.s.l.; 12–14 May 1974; S. Peck leg.; carrion; DEBU. COSTA RICA – Alajuela • 1 ♀; Volcán Tenorio, N slope near Bijagua Biological Station; 700 m a.s.l.; 18 Jun. 2000; Buck and Marshall leg.; pan traps in tree fall; DEBU • 1 ♀; Río Peñas Blancas; 700 m a.s.l.; 18 Aug. 1986; L. Masner; DEBU • 1 ♂; Florencia Forest; 28 Feb. 1980; H. Howden leg.; dung trap; DEBU. – Cartago • 7 ♂♂, 7 ♀♀; Turrialba, CATIE; 600 m a.s.l.; 26 Feb. 1980; H. and A. Howden leg.; CNCI • 6 ♂♂, 7 ♀♀; Turrialba, CATIE; 600 m a.s.l.; 26 Feb. 1980; H. and A. Howden leg.; INBC • 6 ♂♂, 9 ♀♀; Turrialba, CATIE; 600 m a.s.l.; 28 Feb. 1980; H. and A. Howden leg.; DEBU • 1 ♂; Turrialba, CATIE, Florence Forest; 600 m a.s.l.; 28 Feb. 1980; H. and A. Howden leg.; cup traps; DEBU. – Guanacaste • 1 ♂; Guanacaste Conservation Area, Pitilla Field Station; 29 Jan. 1996; J. Noyes leg.; Malaise; DEBU. – Heredia • 1 ♀; La Selva; 2 Feb. 1990; N. Grieg leg.; around dung ball rolled by Canthon moniliatus Bates, 1887; DEBU. – Limón • 4 ♀♀; Estrella Valley, Pandora; 20 Feb. 1984; H. Howden leg.; carrion trap; CNCI • 1 ♂, 1 ♀; 4 km NE of Bribri; 50 m a.s.l.; Dec.1989 – Mar. 1990; P. Hanson leg.; INBC. CUBA • 1 ♀; Santiago, Gran Piedra Meteorological Radar; 1100 m a.s.l.; 6–17 Dec. 1995; S. Peck leg.; elfin forest, carrion traps; DEBU. ECUADOR – Napo • 2 ♂♂; Tiputini Biodiversity Station, vicinity Yasuní National Park; 0°38′ S, 76°0′ W; 14–19 Feb. 1998; D.C. Darling leg.; human dung pitfalls; DEBU • 1 ♀; Yasuní National Park, Yasuní Research Station; 0°38′ S, 76°36′ W; 3–20 Nov. 1998; Pape and Viklund leg.; rain forest, Malaise trap; DEBU • 4 ♀♀; Jatun Sacha Reserve, 6 km E of Misahuallí; 1°04′ S, 77°37′ W; 450 m a.s.l.; 30 Apr.–7 May 2002; Buck et al. leg.; landslide in forest, Malaise trap; QCAZ • 2 ♂♂; Tena; 500 m a.s.l.; 21–27 May 1987; Brown and Coote leg.; secondary rainforest, Malaise head; DEBU • 3 ♂♂; 12 km SW of Tena; 500 m a.s.l.; 8–11 Aug. 1976; S. Peck leg.; dung trap; QCAZ • 1 ♂; S side of Rio Piocullin, SW Puerto Napo, S of Limonchicta; 600 m a.s.l.; 23–27 May 1987; primary lowland rainforest, Malaise head; ROME. – Pichincha • 8 ♂♂, 8 ♀♀; Rio Palenque Station, 47 km S of Santo Domingo; 250 m a.s.l.; 17–25 Feb. 1979; S.A. Marshall leg.; dung; DEBU • 7 ♂♂, 3 ♀♀; Rio Palenque Station, 47 km S of Santo Domingo; 250 m a.s.l.; 17–25 Feb. 1979; S.A. Marshall leg.; QCAZ • 1 ♂; Rio Palenque; 25 Feb. 1979; S.A. Marshall leg.; carrion; QCAZ • 1 ♀; Rio Palenque; 27 Feb. 1979; S.A. Marshall leg.; carrion; DEBU • 1 ♂; Palenque; 24–25 Feb. 1975; S. Peck leg.; day 3, trap 35; DEBU • 13 ♂♂, 23 ♀♀; Rio Palenque; 25 Feb. 1979; S.A. Marshall leg.; dung; DEBU • 4 ♂♂, 17 ♀♀; Rio Palenque; 27 Feb. 1979; S.A. Marshall leg.; dung; DEBU • 5 ♂♂, 17 ♀♀; Rio Palenque; 27 Feb. 1979; S.A. Marshall leg.; dung; QCAZ • 1 ♂; Rio Palenque; 22–23 Feb. 1976; S. Peck leg.; dung trap; DEBU • 1 ♂; Rio Palenque; 25–26 Feb. 1976; S. Peck leg.; dung trap; DEBU • 1 ♂; Rio Palenque; 22 Feb. 1976; S. Peck leg.; DEBU. GUATEMALA – Baja Verapaz • 2 ♂♂, 2 ♀♀; 6.6 km W of Chilasco; 1700 m a.s.l.; 30 May 1991; H. Howden leg.; dung; DEBU. – Guatemala • 1 ♂, 1 ♀; Santa Catarina Pinula; 1840 m a.s.l.; 11–13 Jun. 1991; B.D. Gill leg.; dung traps; DEBU. – Petén • 11 ♂♂, 7 ♀♀; Tikal; 28–30 Jul. 1978; Helava and Kukal leg.; dung trap; DEBU. – Sacatepéquez • 1 ♀; 5 km SE of Antigua; 14°32′14″ N, 90°41′41″ W; 2125 m a.s.l.; 10–13 Jun. 2009; hardwood forest, Malaise trap; DEBU. – Zacapa • 1 ♂; 3.5 km SE of La Unión; 1500 m a.s.l.; 26 Jun. 1993; R. Anderson leg.; cloud forest litter; DEBU • 3 ♀♀; same collection data as for preceding; 25–27 Jun. 1993; Ashe and Brooks leg.; FIT #128; DEBU. GUYANA – Mazaruni-Potaro • 1 ♀; Takutu Mountains; 6°15′ N, 58°55′ W; 8–10 Dec. 1983; Perkins and Steiner leg.; window trap in montane rainforest near logging area; DEBU. – Potaro-Siparuni • 1 ♀; Mount Wokomung; 5°06′35″ N, 59°49′15″ W; 1234 m a.s.l.; 27 Oct.–1 Nov. 2004; B. Hubley leg.; primary rainforest, human dung, pitfall trap; ROME. – Rupununi • 3 ♂♂, 4 ♀♀; Kurupukari, Essequibo River; 200 ft a.s.l.; 9 Oct. 1990; B. Hubley leg.; primary forest, dung traps; ROME • 1 ♂, 1 ♀; Kabocalli, Iwokrama Forest Res.; 60 m a.s.l.; 3–5 Jun. 2001; Brooks and Falin leg.; FIT; DEBU. HONDURAS • 1 ♀; Olancho; 23 May 1995; R. Cordire leg.; FIT; DEBU. MEXICO – Campeche • 3 ♂♂, 1 ♀; Chicanna, 10 km W of Xbuzi; 300 m a.s.l.; 12–14 Jul. 1983; S. and J. Peck leg.; tropical seasonal forest, carrion traps; CNCI • 4 ♂♂, 2 ♀♀; Chicanna, 10 km W of Xbuzi; 300 m a.s.l.; 12–14 Jul. 1983; S. and J. Peck leg.; tropical seasonal forest, carrion traps; UNAM • 3 ♂♂, 7 ♀♀; 53 mi. E of Escárcega; 500 ft a.s.l.; 8–14 Aug. 1971; A. Newton leg.; tropical semi-evergreen, dung; DEBU • 3 ♂♂, 7 ♀♀; same collection data as for preceding; FMNH • 7 ♂♂, 9 ♀♀; 87 mi. E of Escárcega; 800 ft a.s.l.; 8–14 Aug. 1971; A. Newton leg.; semi-evergreen, human dung; DEBU • 10 ♂♂, 14 ♀♀; same collection data as for preceding; UNAM • 2 ♂♂, 2 ♀♀; same collection data as for preceding; FMNH. – Chiapas • 2 ♀♀; 21 mi. N of Bochil; 5500 ft a.s.l.; 18–24 Aug. 1971; A. Newton leg.; pine, oak, Liquidambar L., human dung; DEBU • 1 ♀; 4 km SE of Custepec; 15°42′30″ N, 92°55′51″ W; 2100 m a.s.l.; 20 May 2008; cloud forest, Malaise trap; DEBU • 2 ♀♀; Playón de la Gloria; 16°08′53″ N, 90°53′48″ W; 180 m a.s.l.; 25 May 2008; mature wet forest, Malaise trap; UNAM • 1 ♀; Playón de la Gloria; 16°09′37″ N, 90°54′7″ W; 160 m a.s.l.; 25 May–24 Jun. 2008; mature wet forest, Malaise trap; UNAM • 2 ♂♂, 4 ♀♀; 8 km SE of Salto de Agua; 17°30′45″ N, 92°17′40″ W; 60 m a.s.l.; 14–17 Jun. 2008; secondary wet forest, Malaise trap; UNAM • 2 ♀♀; 8 km SE of Salto de Agua; 17°30′58″ N, 92°18′05″ W; 100 m a.s.l.; 14 Jun. 2008; wet forest edge, Malaise trap; UNAM • 5 ♂♂, 8 ♀♀; Palenque; 100 m a.s.l.; 6–9 Jul. 1983; S. and J. Peck leg.; rainforest, carrion trap; DEBU • 1 ♀; Parque Nacional Sumidero; 1000 m a.s.l.; 1 Jun. 1990; H. and A. Howden leg.; CNCI • 1 ♂, 2 ♀♀; 15 mi. NW of Ocozocoautla; 2800 ft a.s.l.; A. Newton leg.; rainforest, dung; DEBU • 5 ♂♂, 2 ♀♀; 11 mi. NW of Ocozocoautla; 3400 ft a.s.l.; 19–25 Aug. 1971; A. Newton leg.; oak-evergreen forest, human dung; FMNH • 6 ♂♂, 4 ♀♀; 4 mi. S of Palenque; 600 ft a.s.l.; 7–15 Aug. 1971; A. Newton leg.; rainforest, human dung; DEBU • 10 ♂♂, 10 ♀♀; same collection data as for preceding but 700 ft a.s.l.; DEBU • 11 ♂♂, 10 ♀♀; same collection data as for preceding; FMNH • 1 ♂; Palenque; 80 m a.s.l.; 2–23 Jul. 1983; S. and J. Peck leg.; secondary vegetation; FIT; DEBU • 1 ♂; Laguna Belgica, 16 km NW of Ocozocoautla; 970 m a.s.l.; 13 Jun. 1990; H. and A. Howden and B. Gill leg.; F.I.T.; DEBU. – Guerrero •

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