Empis (Enoplempis) tripenicillata Sinclair, Brooks & Cumming, 2025, sp. nov.

Empis (Enoplempis) tripenicillata sp. nov. urn:lsid:zoobank.org:act: FC1F8351-E511-483F-9D69-50E90A0715CE (Figs 167–169, 172, 173, 175) Type material. HOLOTYPE ♂ (Fig. 167), labelled: “USA: WA: Lewis Co., trib./ of Catt Creek, 2847 ft,/ 46°41′53″N 121°57′46″W,/ 16.vi.2018, S.E. Brooks,/ CNC1109361 ” “ HOLOTYPE / Empis (Enoplempis) / tripenicillata Sinclair,/ Brooks & Cumming [red label]” (CNC). PARATYPES: CANADA. British Columbia: Agassiz [49°14′N 121°46′W], 7.v.1927, R. Glendenning (1♂, CNC); Horne Lk. [49°20′N 124°42′W], 4.vi.1955, R. Coyles (1♂, CNC); Qualicum Bay [49.398°N 124.613°W], 29.v.1955, R. Coyles (1♂, CNC). USA. California: Del Norte County: Six Rivers NF, Fire Rd 17N21 nr Myrtle Ck, 41.870°N 124.040°W, 1.vi.2009, K. Collins & I.S. Winkler, E09-41 (1♂, CNC). Mendocino County: NCCRP, 3 mi. N Branscomb [39°42′N 123°39′W], 1400 ft, 17.v.1975, S. Mikasaka (1♂, EMEC); same data except, 30.v–1.vi.1980, N. Davidson (1♂, EMEC); same data except, J. R. Wilson (1♂, EMEC). Sonoma County: Cazadero [38°32′N 123°5′W], 12–14.iv.1918, Cornell Univ. Lot 542, Sub, J.C. Bradley (1♂, CUIC). Oregon: Benton County: Corvallis, Lewisburg Saddle, below Old Growth Tr., 44°38′32.28″N 123°17′20.76″W, MT over creek, 23.iv–6.v.2013, SJF (1♂, CNC). Clackamas County: Mt Hood NF, Tollgate Cpgd [45°19′N 121°54′W], T3 S R7 E Sec. 11, 22.vi.1977, DDW (1♂, CAS). Coos County: Rogue R, Siskiyou NF, NFD 5325, ca 3.5 mi. from jct. 33, high dry open woods, 42.675843°N 124.127834°W, 24.v.2014, SJF (1♂, CNC); Rogue R, Siskiyou NF, Coquille R Falls tr. #1257, ca 42.713657°N 124.022767°W, 24.v.2014, SJF (1♂, CNC). Curry County: Rogue R, Siskiyou NF, NF-23 8.6 mi. jct. NF-33, sweep dry woods, 42.627418°N 123.955349°W, 26.v.2014, SJF (1♀, CNC). Douglas County: Glide [43°19′N 123°4′W], 28.iv.1964, K. Goeden (2♂, ODAC); Umpqua NF, 9.5 km NE Tiller, 42°59.16′N 122°51.47′W, 22.vi.2008, C. Borkent, CJB08-17 (1♂, LEM). Hood River County: Hood River [45°42′N 121°31′W], Childs (1♂, USNM); same data except, 1.vii.1917 (4♀, USNM). Josephine County: Grayback Camp [42°08′N 123°27′W], Siskiyou NF, 19.vi.1962, C.A. Toschi (1♂, EMEC). Lane County: N end Blue R Res. [44°10′N 122°19′W], 13.v.1983, D.S. Chandler, sweep (1♂, UNHC); Oakridge, 18 mi. NW [43°52′N 122°42′W], 12.v.1969, E.M. & J.L. Fisher (13♂, 1♀, CSCA); H.J. Andrews Exp. For. [44°13′N 122°10′W], 1450 ft, R. S.7, 18.v.1983, sweep, D.S. Chandler (2♂, UNHC); same data except, R. S.20, 2250 ft, Rd 134, 12.v.1984, flying (1♂, 1♀, UNHC); same data except, Rd 130, 1750 ft, sweep (2♂, 1♀, UNHC); same data except, Lookout Ck, flying (1♂, 1♀, UNHC). Lewis County: same data as holotype (3♂, 7♀, CNC). Skagit County: Rockport SP [48°29′N 121°36′W], 7 mi. E Concrere, 16–17.vi.1971, D.N. Ferro (1♂, WSU); same data except, WJT (2♀, WSU). Diagnosis. This species is distinguished from other species of the E. (En.) delumbis species group by the male hind tibia and swollen hind tarsomere 1 with very long dorsal setae, male mid tibia with very long dorsal setae, male lateral abdominal setae long and pale (Fig. 167), setae on male terminalia long and strong, hypandrial setae longer than hypandrium and phallus with spiculate surface from apex to subapical expansion and pair of narrow lateral lobes. Description. Wing length 5.4–6.0 mm. Male. Similar to E. (En.) floricola sp. nov. except as follows: thorax greyish brown in ground-colour; postpronotal lobe, postalar ridge and usually scutellum yellowish. Pleura without pale margins of katepisternum and anepimeron. Legs slender, yellow, with apical portion of femora and tibia brownish, tarsomeres darker; coxae pale, in contrast to greyish pleura (Fig. 167). Femora with pale ventral setulae, shorter and less dense on fore femur. Fore tibia with 2–3 erect anterodorsal setae, longer than width of tibia; apex with circlet of preapical setae, posterior setae longer; anterior face with short appressed setae. Fore tarsomere 1 inflated, slightly broader than width of tibia, without long dorsal setae, row of strong anterior setae, fine ventral setae and biserial row of slightly stronger setae; tarsomeres 2– 5 with biserial row of strong ventral setae; tarsomeres 1–4 with apicolateral pair of strong setae, longer than width of segment. Mid femur with strong, anterior preapical seta. Mid tibia with row of very long, erect posterodorsal setae, length 3 times width of tibia; ventral and posterior faces with dense fine setulae; circlet of strong preapical setae. Mid tarsomere 1 slender, with long dorsal setae, length twice width of tarsomere; tarsomeres 1–5 with biserial row of strong ventral setae; tarsomeres 1–4 with apicolateral pair of strong setae, longer than width of segment. Hind femur with long dorsal setae on distal half, longer than width of femur; with 2 closely approximated preapical ventral protuberances and third offset protuberance nearly perpendicular to paired protuberances; protuberances with dark, curved, flattened apex or flattened setae; apex with preapical row of 6 flattened, black posterior setae; preapical setae longer than width of femur (Figs 168, 169). Hind tibia with narrow base, without subbasal swelling and bend; basiventral swelling divided into anteroventral and posteroventral setose lobes; clear, thin basal ridge extended to posteroventral lobe; long, numerous dorsal setae, length more than 3 times width of tibia; long anteroventral and posteroventral setae, shorter than dorsal setae (Figs 167–169). Hind tarsomere 1 inflated with long dorsal setae similar to tibia, ventral strong setae longer than width of tarsomere; remaining tarsomeres similar to mid tarsus. Abdomen paler than thorax, with whitish pruinescence; lateral margins of tergites with long pale setae. Pregenital segments modified: tergite 6 with broad U-shaped posterolateral margin. Tergite 7 with posterolateral margin with dark setae; dorsolaterally with knob-like, shiny swelling, flattened dorsally. Tergite 8 with narrow, darkened posterior margin, broadly separated from broader, darkened anterior margin; sclerite fused laterally to sternite 8; posterior margin of ring sclerite with long, dark setae. Terminalia (Figs 172, 173): brown with long, strong dense setae. Cercus short, subtriangular, projecting obliquely from epandrium; apical half slightly tapered; apical setae longer than width of cercus. Subepandrial process extending to either side of phallus. Epandrial lamella subtriangular, tapered, posterior margin rounded with short, upturned projection; setae longer than width of epandrium, stronger near anteroventral margin. Hypandrium long, tapered, apical margin rounded; setae strong, longer than length of hypandrium. Phallus (Fig. 173) with basal half narrow; paired slender lateral processes beyond mid-length; spiculate surface on subapical swelling, extending to broad apex; ejaculatory apodeme half-length of epandrium, Y-shaped, with ventrally positioned lateral apodemes. Female. Similar to male, except as follows: frons broader, slightly wider than anterior ocellus; hindleg without modified setae and processes; hind tibia with long anterodorsal and posterodorsal setae, twice width of tibia; ventral surface of femora as in male; abdominal pleural membrane without darkened, expanded region; tergites 7 and 8 with pruinescence. Geographical distribution and seasonal occurrence (Fig. 175). Empis (En.) tripenicillata sp. nov. is known from southern British Columbia, Washington, Oregon and northern California. Adults have been collected from April to July. Etymology. The species name is Latin for three and painter’s brush, in reference to the row of three short lobes on the hind femur in this species, one of which ends in a setal cluster that looks like a painter’s brush. Nuptial gift presentation. Form unknown.

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