Alloperla elevata Frison 1935

Alloperla elevata Frison (Figs. 9‐16) Alloperla elevata Frison 1935: 335. Holotype male, INHS, Floras Creek, Curry Co., Oregon, examined. Material examined. CANADA: BRITISH COLUMBIA: Chilliwack River, Vedder Crossing, 8 May 1938, S. Spencer, 2 ♂, 4 ♀ (CNIC); Sweltzer Creek, near Cultus Lake, 23 July 1936, W.E. Ricker, 3 ♂, 3 ♀ (BYUC); Vancouver Island, Courtenay, June 1965, N.L.H. Krauss, 1 ♂, 1 ♀ (BYUC). USA: CALIFORNIA: Del Norte Co., Klamath River, Hwy 169, Klamath Glen, 8 June 2005, R.W. Baumann & B.C. Kondratieff, 6 ♂, 1 ♀ (BYUC, CSUC); Smith River, Hwy 199, 3 mi. NE Hiouchi, 7 June 2005, R.W. Baumann & B.C. Kondratieff, 1 ♂ (BYUC). Humboldt Co., Klamath River, Aikens Creek, 17 May 2007, J.J. Lee, 5 ♂, 5 ♀ (BYUC); Mad River, Mad River Fish Hatchery, 5 May 2007, J.J. Lee, 6 ♂, 4 ♀ (BYUC); Mattole River, Camp Gephart, near Petrolia, 19 May 2007, J.J. Lee & G. Gephart, 1 ♂, 1 ♀ (BYUC); Redwood Creek, Hwy 101, 14 May 2005, J.J. Lee, 4 ♂, 2 ♀ (BYUC). OREGON: Benton Co., Alsea River, Hwy 34, Salmonberry Park, 11 June 2005, R.W. Baumann & B.C. Kondratieff, 1 ♂ (BYUC). Curry Co., Floras Creek, 22 May 1933, Dimick & Zella, 1 ♂, holotype (INHS); 20 May 1933, Dimick, 1 ♀, allotype (INHS); Sixes River, Sixes River Campground, 9 June 2005, R.W. Baumann & B.C. Kondratieff, 2 ♂ (CSUC); Winchuck River, Winchuck River Campground, 1 June 1991, R.W. Baumann & B.P. Stark, 4 ♂ (BYUC); 8 June 2005, R.W. Baumann & B.C. Kondratieff, 1 ♂, 20 ♀ (BYUC). Lincoln Co., Alsea River, Hwy 34, Blackberry Campground, 3 June 1991, R.W. Baumann & B.P. Stark, 1 ♂, 3 ♀ (BYUC); Canal Creek, Canal Creek Campground, 11 June 2005, R.W. Baumann & B.C. Kondratieff, 1 ♂ (BYUC); Schooner Creek, Rd 106, 7 mi. E Lincoln City, 11 June 2005, R.W. Baumann & B.C. Kondratieff, 1 ♂ (BYUC). Tillamook Co., Wilson River, Hwy 6, 10 mi. NE Tillamook, 12 June 2005, R.W. Baumann & B.C. Kondratieff, 15 ♂ (BYUC, CSUC). WASHINGTON: Clallam Co., Bogachiel River, Hwy 101, near Forks, 18 June 1967, R.W. Baumann, 5 ♂, 1 ♀ (BYUC); Bogachiel River, Hwy 101, Bogachiel River State Park, 14 June 2005, R.W. Baumann & B.C. Kondratieff, 7 ♂, 11 ♀ (BYUC, CSUC). Cowlitz Co., Kalama River, north of Kalama, 16 June 1968, R.W. Baumann, 1 ♂, 6 ♀ (BYUC). Grays Harbor Co., Humptulips River, Hwy 101, Humptulips, 18 June 1967, R.W. Baumann, 10 ♂, 10 ♀ (BYUC); West Fork Humptulips River, NE Humptulips, 14 June 2005, R.W. Baumann & B.C. Kondratieff, 2 ♂, 3 ♀ (BYUC). Jefferson Co., Queets River, junction Hartzell Creek, Olympic National Park, 14 June 2005, R.W. Baumann & B.C. Kondratieff, 13 ♂, 7 ♀ (BYUC, CSUC). Lewis Co., Chehalis River, Hwy 6, Rainbow Falls State Park, 13 June 2005, R.W. Baumann & B.C. Kondratieff, 2 ♂, 2 ♀ (BYUC, CSUC); Cowlitz River, Hwy 505, near Toledo, 16 June 1968, R.W. Baumann, 7 ♂, 13 ♀ (BYUC). Mason Co., Skokomish River, Hwy 110, 17 June 1967, R.W. Baumann, 3 ♂, 2 ♀ (BYUC). Pacific Co., Naselle River, NE Naselle, 13 June 2005, R.W. Baumann & B.C. Kondratieff, 2 ♂ (BYUC); North Nemah River, 3 mi. SE Nemah, 13 June 2005, R.W. Baumann & B.C. Kondratieff, 15 ♂, 12 ♀ (BYUC, CSUC). Skagit Co., Sky Creek, Hwy 20, NW Newhalem, 17 June 2004, R.W. Baumann & B.P. Stark, 16 ♂, 4 ♀ (BYUC). Thurston Co., Deschutes River, Hwy 507, near Rainier, 12 June 1968, R.W. Baumann, 1 ♂, 2 ♀ (BYUC); Nisqually River, Nisqually, 13 June 1967, R. W. Baumann, 3 ♂ (BYUC). Male. Macropterous. Length of forewing 9‐11 mm. General color, lime green in life, white in alcohol. Head and pronotum without black markings. Dorsal aspect of epiproct elongate, constricted near middle, apex broadly rounded, with numerous shallow marginal crenulations fringing apex (Figs.11‐14); surface bare in apical half, clothed with dense appressed setae in basal half (Figs. 9‐16); in lateral view, disc shaped (Figs. 9, 10, 15, 16). Stem clothed with fine setae (Figs. 9, 11, 13). Female. Length of wings 10‐11 mm. Coloration and markings similar to male. Subgenital plate produced with posterior margin acute to slightly rounded, general outline triangular, margin usually reaching beyond sternum 9 (Frison 1935, Plate XIV, Fig. 41). Diagnosis. The male of A. elevata can be easily distinguished from both A. severa and A. thalia. by the hour glass shape of the epiproct in dorsal view, enlarged at base, and broadly rounded at apex, bearing fringe of shallow crenulations (Figs. 11‐14) and disc shaped in lateral view (Figs. 9, 10, 15, 16). The subgenital plate of the female of A. elevata is more rounded at the apex (Frison 1935, Plate XIV, Fig. 41) than either A. severa (Lyon and Stark 1997, Fig. 43), or A. thalia (Fig. 24). Remarks. Alloperla elevata is a species that appears restricted to larger Coastal streams and rivers of the Pacific Northwest, from northern California to British Columbia. Ricker (1943) in his work on the stoneflies of southwestern British Columbia considered A. elevata “…enormously abundant on Sweltzer creek, and moderately so in other medium to large streams near sea level. It has been taken in every month but December and January…”. Ricker (1943) described the epiproct of A. elevata as “dumbbell‐ shaped,” however, in 1954 he considered that “straight and dumbbell shapes intergrade, and even coastal specimens vary somewhat in this respect.” Our examination of material indicates that A. elevata, Ricker’s “coastal form”, is consistent in epiproct shape and form (Figs. 9‐16) throughout its known range. Jewett’s (1959) description of the epiproct of A. severa as “dumbbell shaped” and his use of Frison’s (1935) illustration of that species indicates that he might have only examined specimens of A. elevata. Jewett (1960) did not report this species from California. Gaufin et al. (1972) listed Montana records for A. elevata, but did not include it in the key to the species. These specimens could be either A. severa or A. thalia, since both occur in Montana.

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