Anthomyza occidentalis Roháćek & Barber 2016, sp. nov.

Anthomyza occidentalis sp. nov. (Figs 337, 339, 343–356, 377) Type material. HOLOTYPE: ♂, “ USA: OR: Lane Co., Tokatee Klootchman St.Nat.Site., ~7.9kmS Yachats, 10.vi.2009, KN Barber, sweeps, mostly E. tel-mateia, Angelica, base of cliff 44°12.47’N 124°06.90’W ” and “ Holotypus ♂ Anthomyza occidentalis sp. n., J. Roháček & K. N. Barber det. 2014” (red). The specimen (largely yellow coloured, Fig. 339) is in good condition, with partly visible gonostylus (DEBU, intact). PARATYPES: CANADA: BRITISH COLUMBIA: Milner, 12.vii.1953, 1 ♀, G. J. Spencer leg. (CNCI, genit.prep., right wing missing). UNITED STATES OF AMERICA: CALIFORNIA: Cambria, Santa Rosa Creek Trail, 35°33.94'N 121°06.03'W, sweeps, Equisetum telmateia braunii & Delairea odorata under canopy, 3.v.2014, 1 ♀, 4.v.2014, 1 ♂, K. N. Barber leg. (LACM); Mendocino Co., Inglenook Fen, fen area, 30–50', 11.viii.1972, 2 ♂♂ 1 ♀ (1 ♂ 1 ♀ genit. prep., 1 ♂ with det. as Anthomyza pallida), 22.vii.1972, 2 ♀♀ (1 ♀ genit. prep.), E. I. Schlinger leg. (EMEC, UCIS- 215194, -233, -249, -597, -599); Marin Co., 2 air mi W. Inverness, 1.v.1976, 1 ♀, J. Doyen & P. Rude leg. (EMEC, genit. prep.); San Francisco Co., Lake Merced, 7.v.1927, 1 ♂, C. L. Fox leg. (genit. prep., wing illustration); San Francisco, Lake Merced, collected with night trap in willow thicket, 21.vi.1964, 2 ♂♂ (both genit. prep.), collected in willow thicket near lake, 1.viii.1964, 3 ♂♂ (2 ♂♂ genit. prep.), P. H. Arnaud Jr. leg.; Humboldt Co., McKinleyville bog area nr. Azalea Avenue, 9.vii.1980, 1 ♂ 1 ♀ (1 ♂ genit. prep.), T. W. Davies leg. (all CASC); Morro Bay, 27.vii.1940, 1 ♂ (genit. prep.), A. L. Melander leg.; Pacinc Grove, 4.vii.1921, 1 ♂, A. H. Sturtevant leg. (both USNM); Marin Co., Point Reyes, 19.iv.1980, 1 ♂ 1 ♀, S. A. Marshall leg. (DEBU); Humboldt Co., Prairie Ck. Redwoods S. P., Elk Prairie Cmpgd., 41°21.63'N 124°01.73'W, sweeps, riparian Scirpus sp., 8.vi.2009, 1 ♂, K. N. Barber leg. (CNCI); Rosemead, 17.iii.1950, 1 ♂ 1 ♀, A. H. Sturtevant leg. (USNM); Los Angeles Co., Rosemead, 17.iii.1950, 5 ♂♂ 4 ♀♀ (3 ♂♂ 1 ♀ genit. prep.), 22.iii.1950, 1 ♂ 1 ♀ (1 ♂ genit. prep.); San Mateo Co., San Francisco, 18.v.1950, 2 ♂♂ 2 ♀♀ (1 ♂ 1 ♀ genit. prep.), all M. R. Wheeler leg. (all AMNH); San Francisco, 1.viii.1915, 1 ♂, A. L. Melander leg. (USNM, genit. prep.). OREGON: Curry Co., Cape Blanco, 29.vi.1972, 1 ♂, W. N. Mathis leg.; Curry Co., 8 mi N Gold Beach, 29.vi.1972, 3 ♂♂ 1 ♀ (2 ♂♂ 1 ♀ genit. prep.), G. Steyskal leg. (all USNM); Curry Co., Samuel Boardman S. P., Lone Ranch Beach, 42°06.07'N 124°20.76'W, sweeps, mostly Equisetum telmateia, 3.vi.2009, 2 ♂♂; Lane Co., Tokatee Klootchman St[ate] Nat[ural] Site., ~ 7.9 km S Yachats, 44°12.47'N 124°06.90'W, sweeps, mostly E. telmateia, Angelica, base of cliff, 10.vi.2009, 5 ♂♂ 3 ♀♀, all K. N. Barber leg. (CNCI 4 ♂♂ 2 ♀♀, 1 ♂ 1 ♀ genit. prep., SMOC 1 ♂ 1 ♀, both genit. prep.). WASHINGTON: Friday Harbor, 6.vii.1905, 1 ♂ 2 ♀♀, J. M.Aldrich leg. (USNM, 1 ♂ 1 ♀ genit. prep., 1 ♀ with det. as Anthomyza pallida Zett.); Pierce Co., Tacoma, 22.vi.1982, 1 ♀, T. L. Whitworth leg. (LACM); Whidbey Island nr. Keystone Ferry, 18.ix.1975, 1 ♀, G. F. Hevel leg. (USNM). Other material examined (not included in type series). UNITED STATES OF AMERICA: CALIFORNIA: Asilomar, 1.ix.1945, 1 ♂, A. L. Melander leg. (headless, genit. prep.); Pacinc Grove, 4.vii.1921, 1 ♂ (headless, genit. prep.), A. H. Sturtevant leg. (both USNM). OREGON: Curry Co., 8 mi N Gold Beach, 29.vi.1972, 1 ♂, G. Steyskal leg. (USNM, headless, genit. prep.). WASHINGTON: Whidbey Island nr. Keystone Ferry, 18.ix.1975, 1 ♀, G. F. Hevel leg. (USNM, genit. prep., legs and one wing missing). Description. Male. Total body length 2.58–3.10 mm. Externally very similar to A. concolor including the (slightly less) variable body colour (Fig. 339), ranging from largely yellow to largely brown, although the latter variant seems to be comparatively infrequent. Head about as long as high or somewhat longer than high, anteriorly distinctly angular in pronle and face receding (more than in A. concolor), largely yellow but frons and particularly occiput with variable pale to dark brown markings. Occiput distinctly concave medially, in pale specimens light yellow with dark yellow to ochreous brown crescent-shaped areas surrounding the medial pair of silvery white microtomentose spots; in darker specimens these areas darker and widened laterally; in darkest specimens occiput largely brown to dark brown (including small spot behind ocellar triangle), with only medial area above foramen (= ground of silvery white microtomentose spots) yellow and connected dorsally with orbital stripes. Frons relatively narrow, largely dull, pale to dark yellow with only ocellar triangle brownish (lighter in pale specimens), in darkest adults also middle of frontal triangle (usually pale) brown and ocellar triangle blackish brown; frontal triangle subshining, with sparse whitish grey to white microtomentum (denser and nner on ocellar triangle). Orbits pale to whitish yellow with sparse silvery white microtomentum as in A. concolor. Frontal triangle narrower (posteriorly only slightly wider than ocellar triangle) and shorter than in A. concolor, reaching to anterior third of frons. Frontal lunule small, yellow. Face, parafacialia and gena formed, coloured and microtomentose as in A. concolor; postgena, ventral part of occiput and mouthparts pale yellow to yellow (also in darkest specimens). Cephalic chaetotaxy (all setae black) as in A. concolor but pvt usually (sometimes surprisingly) long and strongly crossed, no additional microsetula in front of shorter anterior ors setula; subvibrissa similar to foremost peristomal setula and peristomals (4–6) usually shorter. Palpus as in A. concolor, including chaetotaxy. Eye subovoid, broader anteroventrally, with longest diameter oblique and 1.5–1.6 times as long as shortest. Gena lower than in A. concolor, with shortest height 0.11–0.13 times as long as shortest eye diameter. Antenna geniculate, entirely yellow; 1st nagellomere with white pilosity somewhat longer than in A. concolor. Arista with basal segments ochreous to brown and distal setiform part blackish brown, the latter about 2.1 times as long as antenna, shortciliate (cilia somewhat shorter than those on 1st nagellomere). Thorax hardly narrower than head. Scutum with variable colouration ranging from almost entirely yellow, through yellow with various brownish darkened areas (lateral stripes outside of dc lines, medial area in posterior half) to almost entirely dark brown (usually with paler brown notopleural area). Scutellum yellow, with various brown darkening medially to almost completely dark brown (often with paler margins). Dorsum of thorax with distinct yellowish grey microtomentum (best visible in darkest specimens) and relatively dull. Pleural part of thorax more shining than scutum, most often entirely yellow or with various brown markings (more dorsally) to largely dark brown with only small areas (usually ventrally) ochreous brown to yellow. Postscutellum (this always darker) and postnotum yellow to ochreous to dark brown. Thoracic chaetotaxy very similar to that of A. concolor but anterior dc longer than all other setae on scutum except for posterior dc (being longest thoracic seta) and hindmost dc microseta unusually enlarged and resembling a third dc macroseta (sometimes there is 1 additional, more anterior, enlarged dc microseta); sternopleuron with 4–6 upcurved setulae in dorsal half and with 4–5 longer setae ventrally. Scutellum rounded subtriangular, slightly to distinctly convex dorsally. Legs pale yellow to deep yellow (also in darkest specimens in contrast to A. concolor), only distal half to four-nfths of last tarsal segment of all tarsi dark brown. Pedal chaetotaxies as in A. concolor but f 1 with ctenidial spine as long as or slightly longer than maximum width of t 1 and f 3 with 6–9 (usually 8–9) shortened and thickened setae in posteroventral row. Wing (Fig. 337) extremely similar to that of A. concolor including venation but R 4+5 and M usually more distinctly bent and r-m often situated distinctly in front of middle of discal cell (dm). Wing measurements: length 2.79–3.38 mm, width 0.97–1.12 mm, Cs 3: Cs 4 = 0.97–1.12, rm\dm-cu: dm-cu = 2.15–2.61. Haltere yellowish white with slightly darker stem (as in A. concolor). Abdomen also variable in colouration, although less so than in A. concolor. Colour of preabdominal terga (T1–T5) largely yellow to ochreous only in palest specimens and completely brown in darkest specimens, but in others with various darker markings: T1–T3 usually more darkened than T4–T5 or T1–T3 entirely brown and T4–T5 partly ochreous yellow. T1–T5 sparsely greyish microtomentose and subshining, with relatively short and sparse setae as in A. concolor. T1 and T2 dorsally distinctly separate but laterally fused. T1 shortest, T2 slightly shorter than T3–T4, the latter subequal in size; T5 slightly longer than T4, all terga bent onto lateroventral sides of abdomen. Preabdominal sterna pale yellow to ochreous, in dark specimens S1, S2 and S5 (this usually partly) brown darkened, all relatively broad and becoming slightly wider posteriorly; S1 as in A. concolor, S2 slightly wider than long, S3–S5 about 1.5 times wider than long; S3–S4 transversely suboblong, S5 not wider than S4 but with posterior margin slightly emarginate. S2–S5 nnely but not densely setose, S1 bare and with usual darker posterior stripe. T6 very short and transverse, bare, submembranous, pale yellow to ochreous brown but with only lateral parts pigmented (usually poorly visible). S6–S8 brown to dark brown, S8 darkest but usually paler than epandrium. S6 and S7 with darker anterior marginal ledge, S6 with 2–3, S7 with 2 relatively long setae; S8 about as long as epandrium, paler at anterior margin, setose in posterior half. Genitalia. Epandrium (Figs 343, 344) uniformly brown to blackish brown, distinctly longer, of shorter height and part

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