Trimma ukkriti Winterbottom 2021, new species

Trimma ukkriti new species Ukkrit’s Pygmygoby Figs. 1–4. Trimma okinawae (non Aoyagi 1949:173): Winterbottom et al. 2014:98 (DNA barcode analysis, in part, Thailand specimens) Material examined. A total of 73 specimens (8 of which are tissues) from the seas around Phuket, Thailand (Andaman Sea, Malacca Strait). The samples were collected by Richard Winterbottom, Wouter Holleman, Randy Mooi, and Ukkrit Satapoomin. Holotype: ROM 68764 (23.5), Ko Racha Yai, SE side, patch reef, 7.587222, 98.368611, very varied cor-als— Acropora, Porites, Pectinia, Pavona, Astreopora, Millepora, etc., visibility 10.5 m, 3.0– 6.1 m, rotenone, RW 93– 21, 19 Nov., 1993. Paratypes: PMBC 11448, 11(14.9–26.0), Ko Dok Mai, west side, 30 m N and nearer shore than RW93-27, reef profile dropping away at average of 45°, some caves, 7.797222, 98.534167, some corals, lots of Diadema, whip corals, large siliceous sponges, visibility 10.5 m, 0–15.2 m, rotenone, RW 93–28, 24 Nov., 1993. ROM 68093, (25.5), Ko Aeo, NE side, base of fringing reef towards shore, 7.763055, 98.34, 50% live vs dead coral (Acropora, Porites, Pectinia, Pavona), very little algae, heavy siltation on rubble (mostly staghorn), visibility 4.5 m; 0–6.1m, rotenone, field # RW93–04, 10 Nov., 1993. ROM 68094, 5(10.8–24.6), Ko Racha Yai, S tip, steep (45°) wall of large boulders (up to 3 m diameter), 7.583333, 98.3625, little coral growth or algae, visibility 9 m, 15.2–21.3 m, rotenone, field # RW93–20, 19 Nov., 1993. ROM 68095, 6(10.3–25.9), Ko Hi, bay on S coast about middle of its length, base of fringing reef and adjacent sand/silt interface, 7.741667, 98.375833, 90% live coral, with encrusting Astreopora, Pavona, favids, Porites, Acropora, little algae, visibility 9 m; 6.1-10.1 m, rotenone, field # RW93–05, 11 Nov., 1993. ROM 68756, 2(22.8–23.8), Ko Lon, North side, outer fringing reef off sandy beach, from base of reef landward, 7.786111, 98.395833, 50 % live/dead coral (Acropora, Pavona, Porites bommie), little algae, heavily silted, 0–9.1 m, rotenone, field # RW93–03, 10 Nov., 1993. ROM 68757, 3(10.5–18.0), Ko Hi, bay on S coast about middle of its length, 75 m W of RW93-05, fringing reef, 7.741667, 98.375556, 90 % live coral (Astreopora, Pavona, favids, Porites, Acropora), little algae, visibility 9 m, 6.1–11.0 m, rotenone, field # RW 93–06, 12 Nov., 1993. ROM 68758, 2(18.0–22.4), Ko Hi, bay on S coast about middle of its length, 75 m N of RW93–06, isolated patch reefs with sand, 7.741944, 98.375556, different corals, 50–50 live/dead coral, several stands of Acropora staghorn, visibility 6 m; 4.9 m, rotenone, field # RW93–07, 12 Nov., 1993. ROM 68759, 2(14.5–16.1), Patong Beach, off Sandy Beach near north tip of bay, coral slope (45°) to flat silt bottom, 7.92083, 98.27111, 80-90% live coral (Pavona, Astreopora, encrusting corals, favids, etc.), visibility 7.6 m, 7.6–12.8 m, rotenone, field # RW 93–08, 13 Nov, 1993. ROM 68760, 2(17.8–18.0), Kata Bay, W side, 20 m E of small island named Ko Pu, 7.806944, 98.292222, large Porites bommie surrounded by some live (Pavona, Pectinia) coral, visibility 6 m, 3.0– 8.2 m, rotenone, field # RW 93–09, 14 Nov., 1993. ROM 68761, (20.7), Ko Racha Yai, E coast near N tip, heavy patch reef, 7.604167, 98.380556, Porites, Acropora (staghorn), a little silt, visibility 12 m, 3.0– 7.6 m, rotenone, RW 93-12, 16 Nov., 1993. ROM 68762, 10(16.4–26.5), Ko Racha Yai, bay in middle of N coast, 250 m from beach and 100 m from E arm of bay, beach rock arch with live coral, sand/patch reef, 7.607778, 98.368611, Acropora, Porites, Pectinia coral, Millepora, visibility 9 m, 6.1–10.7 m, rotenone, field # RW 93–15, 17 Nov., 1993. ROM 68763, 5(18.5–21.5), Ko Racha Noi, SE end, 200 m N of RW93-17 (off last beach before short break to next island), huge (4.6 m diameter) beach rock bommie surrounded by sand and patch reef, 7.470833, 98.326389, Pectinia, Pavona, Acropora, Pocillopora, visibility 12 m, 1.5–9.1 m, rotenone, RW 93–18, 18 Nov., 1993. ROM 69250, (20.3), Ko Nipit (Nipit Island), north coast, 6.485556, 99.296667, 6 m, Tea Seed extract, 12 Dec., 1994, coll: U. Satapoomin. Tissues: ROM T 00471, (17.7), collected with ROM 68761. ROM T 03797 – T 03803, 7, just off Bangthao Beach, 7.98721, 98.28465, 2008, coll: U. Satapoomin. Non-types: MPM 31145, 13 (10.1–23.8), Ko Mai Thon, NW tip, patch reef, 7.761111, 98.479167, wide variety of corals, visibility 7.6 m, 4.6–8.5 m, rotenone, RW 93–25, 23 Nov., 1993. Diagnosis. A species of Trimma with second spine of first dorsal fin reaching posteriorly to between base of spine and base of 4 th ray of the second dorsal fin when adpressed, at least some branched pectoral fin-rays, fifth pelvic fin-ray unbranched, scales present in predorsal midline, no cheek or opercular scales, 15 or more total gill rakers, cheek with 6 papillae in row c, and a bony interorbital of less than 55% the width of the pupil, posterodorsal interorbital trench not extending laterally beyond 5 th papilla of row p. Cheek and anterior part of head with orange to red spots (light unpigmented areas in preserved specimens) less than pupil-diameter in size, a diamond-mesh pattern formed by melanophores thickly lining entire scale pockets on nape and body. Description. The description is based on the holotype and up to 19 paratypes selected from lots ROM 068095, ROM 69250, ROM 068093, ROM 68094, ROM 68762, ROM 68764, and ROM 68763. Dorsal fin VI + I 8– 9 (9.0), second spine of first dorsal fin reaching to between base of dorsal spine and 4 th fin-ray of second dorsal-fin (Fig. 1), third dorsal spine reaching posteriorly to between inter-dorsal space to base of spine of second dorsal fin, first ray of second dorsal fin branched (75%) or unbranched, remaining fin rays branched except for posterior element of last ray, fin reaches posteriorly 56– 64 –85% (64.13%) distance between base of last ray and first exposed dorsal procurrent caudal-fin ray; anal fin I 9, first ray unbranched in 17 of 20, fin reaching posteriorly 47– 60.1 –70% (60.1%) distance between base of last ray and first exposed ventral procurrent caudal-fin ray; pectoral fin 19– 20 (19.4) with 3– 4 –6 dorsal and 4– 7 –9 ventral unbranched rays and 6– 9 –13 branched rays in central portion of fin, fin reaching posteriorly to vertical above urogenital papilla to base of anal spine; pelvic fin I 5, fifth ray unbranched and 48– 56 –67% (52.4%, 18) length of fourth ray, which reaches posteriorly to between base of anal spine to 4 th anal fin-ray, pelvic rays 1–4 with single sequential branch point; basal membrane either attached to sides of belly just lateral to midline or forming narrow fold across midline dorsal to last pre-pelvic scale; no fraenum. Lateral scales 23 –24 (23.1); anterior transverse scales 10 –11 (10.1); posterior transverse scales 8– 9 (9.0); cheek and opercle scaleless; predorsal midline crossed by 6– 7 –8 (7.4) irregular scale rows, area in front of first dorsal spine scaleless for about 2 scale-widths but no fleshy crest present, anteriormost extent of scales to just posterior to posterior margin of eye; 3 vertical rows of cycloid scales on pectoral-fin base with 1– 2 scales in anteriormost row, 4– 5 in second row and 5 in outer row; 7 –8 (7.1, 16) cycloid scales in midline anterior to pelvic-fin base; area between pelvic spine and ventral margin of pectoral-fin base with cycloid scales; anterior few rows of scales in midline of belly cycloid; anteriormost row of body scales beneath axil of pectoral-fin base of either cycloid or ctenoid scales. Circumpeduncular scales 12, scale rows in midline between base of last anal ray and first ventral procurrent caudal-fin ray 8– 9 (8.3, 19). Upper jaw with outer row of about 8–10 spaced, enlarged curved canines reaching to distal end of premaxilla, space between adjacent teeth about equal to canine height, gradually decreasing in size posteriorly; several irregular rows of small conical teeth behind outer teeth, gradually decreasing in size and becoming reduced to single row, continuing posteriorly to distal tip of premaxilla; innermost few teeth near symphysis slightly larger and posteriorly oriented. Lower jaw with 4–6 enlarged, spaced, curved canines in outer row reaching to bend of dentary, space between adjacent teeth about equal to canine height; 2–3 irregular rows of slightly curved small teeth at symphysis to dentary bend; innermost row half-length of outer and reaching posteriorly to just beyond bend of dentary, continuing as a single row of small straight conical teeth to dorsal rim of coronoid process. Tongue broadly truncate. Gill opening extending anteroventrally to below mid- to posterior margin of pupil; gill rakers 4 –5 + 13– 14 –15 = 18 –19 (4.1 + 14.0 = 18.1). Anterior naris in short tube reaching anteriorly to above anterior margin of upper lip, posterior opening pore-like with raised rim, separated from bony front of orbit by 1 –1.5 times its diameter, nasal sac raised and on anterior half of snout. Bony interorbital width 26– 27 –37% (31.4%) pupil diameter; moderate U-shaped interorbital trench with no median raised fleshy ridge; narrow, poorly developed postorbital groove ending at papilla 5 of row p (Fig. 2 B); epaxialis reaching anteriorly in midline to vertical above posterior margin of pupil; no dermal ridge in midline of nape extending anteriorly from origin of first dorsal fin although the midline is scaleless for a space of about two scale-widths immediately anterior to first dorsal fin spine. Bony width of interorbital 26– 27 –37 (31.4). Caudal peduncle depth as percentage of caudal peduncle length 41– 42 –52 (45.8); head length as percentage of SL 29– 31 –33 (30.3%); as percentage head length: horizontal eye diameter 33– 35 –40 (36.5%); snout length 19– 24 (22%); cheek depth 20– 30 –31 (25.4%). Cephalic sensory papillae as in Fig. 2. Number of papillae in each row: a = 6; b = 6– 9 –10 (8.0); c = 6; cp = 1; d = 7– 11 (8.9); dʹ = 7– 12 –14 (10.1); e-anterior = 14– 15 –21 (16.6); e-posterior = 16– 20 –24 (

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