Hydrophilus (Hydrophilus) triangularis Say. A 1823

Hydrophilus (Hydrophilus) triangularis Say, 1823 (Figs. 2, 6, 7) Hydrophilus triangularis Say, 1823: 201. Stethoxus (s. str.) triangularis (Say); Bedel 1891: 313. Hydrous (Pagipherus) triangularis (Say); Kuwert 1893: 89. Hydrous (s. str.) triangularis (Say); Zaitzev 1908: 366. Stethoxus subsulcatus LeConte, 1854: 221. Synonymy: LeConte 1855: 367; Knisch 1924: 258. Hydrous (Pagipherus) triangularis var. subsulcatus (LeConte); Kuwert 1893: 89. Hydrophilus trisitis Motschulsky, 1859: 174. Synonymy: Gemminger and Harold 1868: 476. Material Examined. See Appendix 1. Diagnosis. Size large (35–38 mm). Specimens often distinctly elongated and narrow relative to co-occurring species. Antennal cupule light in color with no to moderate elongation, ca. 1.5–2x as long as wide and not entirely covering first antennal club segment. Protarsal claws of males unequal in both size and shape. Terminal segment of protarsi greatly widened with outer margin forming an obtuse angle (Fig. 2). The abdominal pubescence forms a distinctive “triangular” lateral margin on the glabrous region on each ventrite (Fig. 6.). This species can be easily separated from other North American species by its elongated shape and very distinctive glabrous abdominal region. Distribution (Fig. 7). CANADA: Alberta, British Columbia, Ontario, Quebec, Saskatchewan; MEXICO: Aguascalientes, Chihuahua, Distrito Federal, Durango, Jalisco, Hidalgo, Michoacán, Morelos, Oaxaca, San Luis Potosí, Tamaulipas; USA: Alabama *, Arizona, Arkansas *, California, Colorado *, Connecticut, Delaware, District of Columbia *, Florida, Georgia, Idaho *, Illinois, Indiana, Iowa *, Kansas *, Kentucky *, Louisiana *, Maryland *, Massachusetts *, Michigan, Mississippi, Missouri *, Montana *, Nebraska, Nevada, New Jersey *, New Mexico *, New York, North Carolina *, North Dakota *, Ohio *, Oklahoma *, Oregon, Pennsylvania, Rhode Island *, South Carolina *, South Dakota *, Tennessee *, Texas, Utah *, Virginia *, Washington, West Virginia *, Wisconsin (Hansen, 1999). Remarks. Localities marked with an asterisk (*) indicate new state records. Hydrophilus triangularis is by far the most widespread and commonly encountered member of the genus in North America. Hydrophilus triangularis CANADA: Ontario: Port Hope, Dr. Bethane (1). Québec: (Literature: Smetana 1988). British Colombia: (Literature: Smetana 1988). Alberta: (Literature: Smetana 1988). Saskatchewan: (Literature: Smetana 1988). MEXICO: Aguascalientes: Aguascalientes, 3-5. viii.1963, P. J. Spangler (1). Chihuahua: Ciudad Jimenez, 26.vi.1964, P. J. Spangler (1). Distrito Federal: “7-8.10” (1); R. Millor (1); Mexico City, Höge (1); Mexico City, O. H. Barrett (1); Mexico City, 28.v.1964, M. W. McFadden (1). Durango: Lerdo Durango, 12.vi.1918, McKinney & Loftin (2). Jalisco: 10 mi N. Chapala, 31.vii.1963, P. J. Spangler (1); 15 mi. N. Chapala, 2.viii.1963, P. J. Spangler (4); Guadalajara, 30.vii.1963, P. J. Spangler (1); 25 mi S. Guadalajara, 6.vii.1964, P. J. Spangler (2); Jocotepec, 11.vii.1959, H. E. Evans (CUIC: 1). Hidalgo: Ixmiquilpan, 1.vii. 1965, P. J. Spangler (2). Michoacán: Zitacuaro, 9.viii.1957, David Lauck (1); Near Cuitzeo, 27. vii.1962, J. R. Zimmerman (1). Morelos: Cuernavaca, xi.1905, Wm. Schaus (1); Cuernavaca, vi (1). Oaxaca: ix.1894, (2); 4 mi SE El Tule, 6.ix.1964, J. R. Zimmerman (1). San Luis Potosí: El Salto, 4.vii.1990, J. Danoff-Burg (SEMC: 1). Tamaulipas: 25 mi. S. Matamoros, 29.viii.1965, P. J. Spangler (11). Unknown: San Antonio, 15.27, L. Stevenson (SEMC: 1). USA: Alabama: Mobile, 10.iv.1949, W. Rosenberg (1); Mobile, 17.iii.1950, W. Rosenberg (1); Mobile, viii.1950, W. Rosenberg (1); Mobile, 19.iii.1949, W. Rosenberg (1); Mobile, 1.iii.1948, W. Rosenberg (4); Mobile, 20.v.1949, W. Rosenberg (1); Jasper, 30.vii.1965, L. G. Sanford (1); “Shefield” [treated as Sheffield], 13.iii.1953, B.D. Valentine (1). Arizona: Arivaca, 3 mi W, 15.vi.1968, O. S. Flint (1); Arivaca, 12.vii.1940, L. C. Kuitert (2); Arivaca, 18.ii.1972, “Lenczy” (1); Arivaca, 11.iv. 1970, “Lenczy” (1); Baboquivari Mountains, 24.vii.1941, Burt Hodgden (1); Green Valley, viii.1972, “Lenczy” (3); 5 mi W Portal, SW Research Station, 8.vi.1956, O. L. Cartwright (1); 5 mi W Portal, SW Research Station, 5.vii.1964, (1); “Chiric Mt”, “3.7”, Hubbard & Schwarz (1); Benson, 15.viii.1970, D. Kichman (1); Kansas Settlement, 0.2 mi W Sulphur Springs Comm. Ctr., 21.vii.1991, Lee Guidry (2); Tucson, Hubbard & Schwarz (1); Tucson, Wickham (1); Tucson, (1); Dos Cabezas, 7.vi.1965, O. S. Flint (1); 11 mi SE Willcox, 6.ix.1969, D. Kichman (1); Tucson, F. H. Snow (SEMC: 2); Tucson, 10.vii.1944, Robert E. Snow (SEMC: 1); Willcox, 24.vii.1970, D. Richman (1); Tanque Verde Mountains, 12.xi.1986, R. S. Beal (1); Bowie, 7.x.1916, E. G. Holt (2); Gila R. at Duncan, 15.iv.1904, F. M. Chamberlain (2); Phoenix, 10-20.viii.1908 (CUIC: 2); Cave Creek, “ 9-10-1917 ”, J. V. M. (1); Kirkland, 24-30.vii.1970, J. May (3); Camp Verde, 22.vii.1969, A. & B. Hilton (3); Montezuma Well, 19.vi.1968, Menke & Flint (1); Winslow, “31-7”, Barber & Schwarz (1); Lake Mary, 20.x.1963, Michael Orr (1); Williams, vi.16, H. A. Kaeber (1); Diamond Creek, Grand Canyon, 8-9.v.1967, D. Davis (1); “ ASU Collection, Arizona” (3); Tucson Mt. Park, iv.1969, R. E. Beer (SEMC: 1); Rt. 170 in Turlingua, 2.viii. 2003, A. E. Z Short (1). Arkansas: Texarkana, 28.viii.1908, E. S. Tucker (1); Hope, Mank Collection (CUIC: 1); Gould, 2.vi.1954, P. H. Thompson (1); Brinkley, 27.ix.1977, Beyers, May, Young (SEMC: 2); Devil’ s Den St. Park, 12.vii.1966, R. W. Hodges (1); nr. Imboden, 10.iv. 1925, B. C. Marshall (1); 19.vii.1968, (1); Yarbro, 17.iv.1966, G. Eikwort (SEMC: 1). California: [without specific locality] (1); Calipatria, 6.vi. 1922, Warwick Benedict (SEMC: 1); Calipatria, 16.vii.1924, Warwick Benedict (SEMC: 1); Laguna Beach (CUIC: 6); Mecca, 11.vii.1923, Warwick Benedict (SEMC: 1); 4.iv.1924, Warwick Benedict (SEMC: 1); Palm Springs, 1932, (1); Los Angeles, 22.viii, C. E. Hutchins (1); Needles, Wickham (1); Needles (1); Oceano, 4.viii.1938, R. I. Sailer (SEMC: 1); Bakersfield, 14.viii.1937, R. E. Beer (SEMC: 1); Bakersfield, 19.viii.1917, Wheeler (1); Panamit Valley, iv.1891, (1), 48 km NNW Baker, 35 41′N, 116 17′W (148 km?), 5.i.1992, W. E. Steiner & J. M. Swearington; Death Valley, iv.1891, K. (1); Fresno, ix, E. A. Schwarz (4); Fresno, 10.vi.1907 (CUIC: 5); Ben Lomond, iv.1931, L. Saylor (1); Bass Lake, 25.viii.1922, Warwick Benedict (SEMC: 1); Travis Air Force Base, 30.ix.1965, Nave & Young (1); Sacramento, 3.v.1921, B. G. Thompson (1); Sacramento, 2.iii. 1921, B. G. Thompson (1); Topaz, “9-4-19”, (1); Colusa, 2.vi.1933, E. C. Zimmerman (1); Willows, 30.viii.1918, A. Wetmore (4); Nr. Seneca, iv.1923, V. S. Barber (1); Pine Creek Valley, 29.vii.2005, G. L. Challet (SEMC: 1); Coquillett (collector) (1); Mink Park, J. Hoyrang (1); Huntington Beach, 7.vi.1937, R. E. Beer (SEMC: 1); El Centro, 26.ix. 1962, Peter D. Ashlock (SEMC: 2). Colorado: Pancha Springs (Treated as Poncha Springs), 14.vii. 1898, E. J. Oslar (1); Buena Vista, 1-6.vii.1896, H. F. Wickham (1); Grand Junction, 29.v.1896, H. K. Plank (1); Denver, Dyar & Caudell (1); Denver, 14.vii.1909, (1); La Porte, 11.vii.1961, J. Toczek (1). Connecticut: Stanford, “ B. T. R. Lab” (treated as Stamford), 1.vii.1929, (1); Stanford, “ B. T. R. Lab” (Treated as Stamford), 19.ix.1929, (1). Delaware: Glasgow, 3.vii.2002, A. E. Z. Short (SEMC: 1). District of Columbia: (3); 1.x.1917, (3); 17.v.1906, W. L. McAtee (1); 1914, M. T. Van Horn (1); 26.vii.1917, (1); 28.iii.1907, D. H. Clemons (5); 27.iv.1904, D. H. Clemons (1); “22.4”, R. P. Currie (5); 3.vi.1902, S. Shoemaker (1); vi.7.1910, (1); vi.10.1910, (1); vii.16.1910 (1); vii.17.1910, (1); vi.1.1902, (1); 4.iv.1910, F. Knab (3); September, F. Knab (1); 4.4.1905, D. H. Clemons (1); 29.iii.1907, W. L. McAtee (1); 24.4.1905, D. H. Clemons (1); “22.4”, H. S. Barber (10); 1.x.1917, (1); 7.ix.1908, D. H. Clemons (1); 27.iii.1907, D. H. Clemons (1); 14.v.1898, C. E. Burden (1); 10.vii.1906, F. Knab (1); 7.ix.1906, D. H. Clemens (3); Hubbard & Schwarz (2); “vi”, Hubbard & Schwarz (1); 24.iv.1895, R. P. Carrie (1); R. P. Carrie (1); 25.iv.1900, H. S. Barber (1); 22.4, R. P. Curio (2); iv.1908, (1); C. E. Burden (1); 5.ix.1902, C. E. Burden (1); 12.vi.1902, C. E. Burden (2); 14.v.1898, C. E. Burden (1); 8.ix.1902, C. E. Burden (1). Florida: Key West, (1); Ramrod Key, 18.viii. 1967, G. F. Hevel (1); Homestead, 17.xi.1951, R. Mansaeti (1); Archbold Biological Station, 15.viii-21.ix.1978, William Rosenberg (3); Charley Apopka Cr at U. S. 17, 14.xi.1979, D. A. Etnier (2); Tampa, Dyar & Caudell (1); Disney World, 21.vi.1979, A. Freidberg (6); 17 mi. N. Tallahassee, Tall Timbers Res. Station, 12-22. vii.1967, L. Collins (1). Georgia: 25.v.1957, W. Rosenberg (1). Idaho: Twin Falls, 29.vi. 1961, T. R. Coupe (1); Boise, 19.v.1925, (1). Illinois: Browns, 27.vi.1956, D. L. Lauck (3); Moose Lake, Albion, 26.vi.1956, D. L. Lauck (1); Wabash R. 1 mi below mouth of Embarras R., 13.ix.1979, Etnier, Rich, Kubb, Bogardus, Clark (1); Bunker Hill, 7 mi. South, 28.v.1953, D. L. Lauck (2); Vandalia, 13.vi.1956, D. L. Lauck (1); Plainview, 19.iv.1957, David Louck (1); Urbana, (1); Urbana, 8.v.1907, Hart & Hood (1); Urbana, viii.1956, D. L. Lauck (1); Chicago, 21.v.1904, W. M. Mann (2); Lily Lake, 11.vii.1956, D. L. Lauck (2); Lake Forest [black square], 13.viii.1899, J. S. Needham (1); Davis, 15.viii.1956, D. L. Lauck (2); Olni Reservoir [possibly “Olney”], 22.v.1953, D. L. Lauck (1). Indiana: (1); Bloomington, 21.vii. 1956, J. R. Zimmerman (1); Bloomington, 22.vii.1956, J. R. Zimmerman (1); Bloomington, 23.v.1954, B. R. Avery (1); Bloomington, 14.iv.1958, J. J. Chopper (1); Bloomington, 24.v.1954, R.D. Curtis (1); Noblesville, 26.vi.1978, H. Jump (4); Grant Plaza, 18.vii.1972 (SEMC: 1). Iowa: Keokuk, May, Shaffer (1); Koekuk, 30.iv.1916, E. Stringham (2); Muscatine, 21.iv.191

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