Lasioglossum (Dialictus) ellisiae

Lasioglossum (Dialictus) ellisiae (Sandhouse, 1924) Figs 18–20, 95B, 99A, 102B, 113B, 120A Halictus (Chloralictus) ellisiae Sandhouse, 1924: 11 (holotype, ♀, deposited in USNM, type no. 26400, USNMENT01570255, examined). Halictus hunteri Crawford, 1932: 72 (holotype, ♀, deposited in USNM, examined) syn. nov. Halictus (Chloralictus) ellisiae – Graenicher 1927: 205 (comparative notes with lepidii). Lasioglossum (Chloralictus) ellisiae – Michener 1951: 1113 (catalogue). Lasioglossum (Chloralictus) hunteri – Michener 1951: 1114 (catalogue). Dialictus tegularis – Mitchell 1960: 423 (in part, synonymy, redescription). — Hurd 1979: 1972 (in part, catalogue). — Moure & Hurd 1987: 134 (in part, catalogue). Lasioglossum tegulare – Donovall & vanEngelsdorp 2010: 16 (in part, checklist). Lasioglossum (Dialictus) tegulare – Krombein 1967: 466 (in part, catalogue). Dialictus hunteri – Hurd 1979: 1966 (catalogue). — Moure & Hurd 1987: 104 (catalogue). Lasioglossum (Dialictus) ellisiae – Gibbs 2009a: 18 (resurrection from synonymy, revision); 2010: 37, 45 (key to species), 120 (redescription); 2011: 25, 32 (key to species), 88 (review). — Wolf & Ascher 2009: 143 (checklist). — Gibbs et al. 2013: 80 (checklist). — Dibble et al. 2017: 13 (checklist). — Kilpatrick et al. 2020: 54 (checklist). Diagnosis Females of L. ellisiae have the tegula somewhat small (reaching and sometimes slightly exceeding posterior margin of mesoscutum in dorsal view), relatively sparsely punctate laterally at midlength (at least IS = 1 PD, up to IS = 1–3 PD), and inner posterior margin weakly concave, with a gentle S-shaped curve coming to a narrow rounded point posteriorly; mesepisternum punctures distinct, round, and about as fine and dense as those of hypoepimeron; metapostnotum shiny with coarse rugae reaching posterior margin; mesoscutum and mesepisternum usually shiny or with weak microsculpture; T1 anterior slope coriarious; T3 without subapical band of tomentum (rarely with some sparse tomentum apicolaterally); and tegula and metasoma sometimes red-orange. Females of L. ellisiae are most similar to those of L. diabolicum sp. nov., L. gaudiale, L. indagator sp. nov., and L. tegulare. Females of L. diabolicum have the tegula slightly smaller (often not reaching posterior margin of mesoscutum in dorsal view) with inner posterior margin usually straight and metapostnotum usually dulled by strong microsculpture. Females of L. gaudiale have the T1 anterior slope shiny with microsculpture absent or very weak and the tegula inner posterior margin more strongly concave. Females of L. indagator have the metapostnotal rugae usually not reaching posterior margin and more extensive tomentum on the mesoscutum lateral and posterior margins, metanotum, and pleura. Females of L. tegulare have the mesoscutum and mesepisternum entirely dull with strong microsculpture and mesepisternum with crowded punctures (IS = 0 PD) some of which are not distinctly round. Some other species in the L. stictaspis species complex (particularly L. stictaspis s. str.) can also be very similar, but have the mesepisternum punctures larger and slightly sparser than those of hypoepimeron, mesoscutum more irregularly and sparsely punctate (as in L. diabolicum), tegula inner posterior margin usually more strongly concave and usually more densely punctate (IS ≤ 1 PD), metapostnotum usually dull with fine rugae, and T3 usually with dense and complete subapical band of tomentum. Males of L. ellisiae have the tegula somewhat small (as in the female); metapostnotum usually shiny with strong subparallel rugae; mesoscutum between parapsidal lines uniformly densely punctate (IS ≤ 1 PD); and discs of T2–T3 becoming more sparsely punctate on premarginal line, not contrasting with impunctate apical rims (punctures in basal half of T2–T3 may be deep and dense (IS ≤ 1 PD) or fine and moderately sparse (IS = 1–2 PD)). Males of L. ellisiae are most similar to those of L. diabolicum sp. nov., L. lepidii, L. puteulanum, and L. tegulare. All of these species have the mesoscutum more irregularly and slightly more sparsely punctate between the parapsidal lines (some IS <1 PD and some IS = 1–2 PD). Males of L. diabolicum and L. lepidii have the disc of T2 uniformly finely and moderately sparsely punctate (IS = 1–2 PD). In direct comparison, males of L. lepidii also have the facial tomentum denser and more tightly appressed (more completely obscuring the surface underneath) and the mesothoracic spiracle more deeply impressed. Males of L. puteulanum and L. tegulare have the disc of T2 uniformly deeply and densely punctate (IS ≤ 1 PD) including the premarginal line. Some other species in the L. stictaspis species complex can also be very similar, but have the tegula slightly larger, with inner posterior margin more strongly concave, and more often entirely densely punctate (IS <1 PD) and mesoscutum more irregularly and sparsely punctate (as above). Etymology Sandhouse (1924) dedicated this species to her fellow bee taxonomist Marion Durbin Ellis. Material examined Holotype UNITED STATES – Massachusetts • 1 ♀; Forest Hills; [42.29° N, 71.11° W]; 5 Aug. 1911; USNM 26400. Other material CANADA – Manitoba • 1 ♀; Delta Beach along Hackberry Road, near Portage La Prairie; 50.18585° N, 98.32591° W; 2–5 Aug. 2007; A.M. Patenaude leg.; WRME • 2 ♀♀; Hadashville; 49.6761° N, 95.8984° W; 28 May 2019; K. Morgan and S. Shukla leg.; WRME 0419384, 0419385 • 1 ♀; Portage La Prairie; 49.93° N, 98.33° W; 27 Jun. 2018; E. Hanuschuk leg.; WRME 0390284. – Ontario • 1 ♀; Haldimand-Norfolk Co., Nixon West Pr.; [42.85° N, 80.4° W]; 11 May 1998; H. Douglas leg.; PCYU • 1 ♀; ibid.; 28 May 1998; H. Douglas leg.; PCYU • 1 ♀; Leamington; [42.05° N, 82.6° W]; 17 Aug. 1987; T. D. Galloway leg.; WRME 0293748 • 1 ♀; Ottawa; [45.42° N, 75.7° W]; 6 Aug. 1981; L. Masner leg.; TAMU-ENTO X1396609 • 1 ♂; ibid.; 6 Aug. 1981; L. Masner leg.; TAMU-ENTO X1383279 • 1 ♀; Pterophylla Plant Nursery; 42.6401° N, 80.573° W; 13–22 Aug. 2006; P.J. Carson leg.; PCYU • 1 ♀; Toronto, Cedarvale Park; 43.69° N, 79.42° W; 30 Aug. 2008; J. Gibbs and Y. Nozoe leg.; WRME 0427965 • 1 ♀; Toronto, Ulster Street; 43.65898° N, 79.41332° W; Oct. 2006; J. Gibbs leg.; PCYU • 1 ♂; Toronto, York campus; 43.773° N, 79.506° W; 11 Aug. 2008; J. Gibbs leg.; WRME 0427964. MEXICO – Tamaulipas • 1 ♀; 8 mi. NE of San Fernando; [24.93° N, 98.06° W]; 27 Aug. 1954; SEMC. UNITED STATES – Colorado • 5 ♀♀; El Paso Co., Foster Ranch, T15 S R65 W Sec. 14 S1/2; [38.741° N, 104.637° W]; elev. 1737 m; 30 May 1976; F.M. Brown leg.; ex Cardaria; UCMC • 2 ♀♀; ibid.; 26 Apr. 1977; F.M. Brown leg.; ex Cymopterus; UCMC • 2 ♀♀; ibid.; 27 Apr. 1977; F.M. Brown leg.; ex Cymopterus; UCMC • 2 ♀♀; ibid.; 3 May 1977; F.M. Brown leg.; ex Cymopterus; UCMC • 1 ♀; ibid.; 31 May 1977; F.M. Brown leg.; ex Cardaria; UCMC • 1 ♀; El Paso Co., Foster Ranch, T15 S R65 W Sec. 14 SW1/4; [38.741° N, 104.642° W]; elev. 1737 m; 29 May 1978; G.H. Kemper leg.; ex Cardaria; UCMC • 3 ♀♀; El Paso Co., Foster Ranch, T15 S R65 W Sec. 22 NE1/4; [38.734° N, 104.651° W]; elev. 1737 m; 10 Jun. 1976; F.M. Brown leg.; ex Cardaria; UCMC • 1 ♀; ibid.; 29 Apr. 1977; F.M. Brown leg.; ex dandelion; UCMC • 3 ♀♀; ibid.; 22 May 1977; F.M. Brown leg.; ex Cardaria; UCMC • 3 ♀♀; ibid.; 22 May 1977; G.H. Kemper leg.; ex Cardaria; UCMC • 1 ♀; ibid.; 31 May 1977; F.M. Brown leg.; ex dandelion; UCMC • 3 ♀♀; ibid.; 31 May 1977; F.M. Brown leg.; ex Cardaria; UCMC • 2 ♀♀; ibid.; 14 Apr. 1978; F.M. Brown leg.; ex Lepidium; UCMC • 2 ♀♀; ibid.; 24 Apr. 1978; F.M. Brown leg.; ex Lepidium; UCMC • 1 ♀; ibid.; 26 Apr. 1978; F.M. Brown leg.; ex Prunus; UCMC • 1 ♀; ibid.; 22 May 1978; F.M. Brown leg.; ex dandelion; UCMC • 2 ♀♀; ibid.; 22 May 1978; F.M. Brown leg.; ex Lepidium; UCMC • 1 ♀; ibid.; 6 Jun. 1978; F.M. Brown leg.; ex Cardaria; UCMC • 1 ♀; ibid.; 25 Jul. 1978; F.M. Brown leg.; ex Melilotus; UCMC • 1 ♀; ibid.; 28 Jul. 1978; F.M. Brown leg.; ex Melilotus; UCMC • 1 ♀; ibid.; 15 Aug. 1978; F.M. Brown leg.; ex Eriogonum effusum; UCMC • 1 ♀; ibid.; 29 Sep. 1978; F.M. Brown leg.; ex Aster; UCMC • 7 ♀♀; ibid.; 10 Jun. 1980; F.M. Brown leg.; ex Cardaria; UCMC • 1 ♀; ibid.; 2 May 1981; F.M. Brown leg.; ex sandy stream bed; UCMC • 1 ♀; ibid.; 22 May 1981; F.M. Brown leg.; ex dandelion; CUIC • 1 ♀; ibid.; 22 May 1981; F.M. Brown leg.; ex dandelion; UCMC • 1 ♀; ibid.; 25 May 1981; F.M. Brown leg.; ex Cardaria; UCMC • 1 ♀; ibid.; 25 May 1981; F.M. Brown leg.; UCMC • 9 ♀♀; El Paso Co., Foster Ranch, T15 S R65 W Sec. 23 N1/2; [38.734° N, 104.638° W]; elev. 1737 m; 13 Jun. 1976; F.M. Brown leg.; ex Cardaria; UCMC • 2 ♀♀; ibid.; 10 May 1977; F.M. Brown leg.; ex dandelion; UCMC • 2 ♀♀; Logan Co.; 40.56755° N, 103.50417° W; 31 May 2012; M. Vandever et al. leg.; CSUC • 1 ♀; Logan Co.; 40.572° N, 103.54303° W; 14 Jun. 2012; M. Vandever et al. leg.; CSUC • 1 ♀; Logan Co.; 40.58675° N, 103.54398° W; 12 Jul. 2012; M. Vandever et al. leg.; CSUC • 1 ♀; Prowers Co., 1 mi. S of Granada; [38.05° N, 102.31° W]; 28 Aug. 1963; N. and B. Marston leg.; AMNH • 1 ♀; Yuma Co., Bonny Dam State Recreation Area, 21 mi. N of Burlington; [39.61° N, 102.18° W]; 10 Jul. 1976; E. R. Hoebeke leg.; CUIC • 1 ♂; Boulder; [40.01° N, 105.27° W]; 1 Aug. 1903; S.A. Rohwer leg.; UCMC • 2 ♀♀; Fort Lupton; [40.08° N, 104.81° W]; 20 Jun. 1950; C.D. Michener leg.; ex Rorippa sinuata; SEMC • 1 ♀; Julesburg; 40.9833° N, 102.25° W; elev. 1055 m; 7 Jun. 1920; AMNH • 1 ♀; Lamar; [38.09° N, 102.62° W]; 22 Jul. 1950; C.D. Michener leg.; ex Melilotus alba; SEMC • 1 ♀; Logan City; 40.58675° N, 103.54398° W; 12 Jul. 2012; M. Vandever et al. leg.; CSUC • 5 ♀♀; Regnier; 37° N, 102.8333° W; elev. 1372 m; 6–9 Jun. 1919; AMNH. – Illinois • 3 ♀♀; Bureau Co., 3 mi. NW of La Moille; [41.56° N, 89.32° W]; 13 May 1970; LaBerge and Molina leg.; ex Salix; INHS • 1 ♀; Champaign Co., Brownfield Woods, Urbana; [40.145° N, 88.165° W]; 29 Mar. 1968; LaBerge and Ribble leg.; INHS • 1 ♀; Kankakee Co., Hooper Branch; [41.02° N, 87.54° W]; 2 Ju

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