Haplostomella multinodosa Kim & Boxshall 2021, sp. nov.
Bibliographic record
Abstract
Haplostomella multinodosa sp. nov. (Figs. 117, 118) Type material. Holotype ♀ (MNHN-IU-2014-21531) and 1 ♀ paratype (MNHN-IU-2014-17392, dissected) from oẚtterella rubra Abbott & Trason, 1968; British Columbia, Canada, seapool rocks (48°49'N, 125°12'15"W), depth 40 m, Pennachetti coll., date unknown. Etymology. The specific name is derived from the Latin mult (=many) and nçd (=swelling), and alludes to the presence of multiple interpodal tubercles associated with legs 1-4. Description of female. Body (Fig. 117A) eruciform, unsegmented, slightly curved dorsally; body length 1.85 mm; maximum width 429 μm (in middle). Anterior part of body tapering anteriorly; cephalosome partially defined by posterolateral margins of cephalic shield. Metasomal region lacking any division, with 5 pairs semicircular dorsolateral folds, each pair located at level of legs 1-5, respectively: last pair of folds smaller than anterior 4 pairs. Genitoabdomen (Fig. 117B) short, occupying less than 20% of body length, unsegmented, not demarcated from metasome, strongly tapering; genital apertures (Fig. 118D) large, positioned dorsolaterally; genital operculum bearing 3 teeth distally and row of 4 denticles on ventral side. Caudal ramus (Fig. 117C) 2.0 times longer than wide (40×20 μm), gradually narrowing distally; armed with 5 setae (1 on outer margin, 1 subdistal dorsal, and 3 distal); seta on outer margin smaller than other caudal setae and positioned at 65% length of ramus. Egg sac (Fig. 117D) about 1.57× 0.41 mm, containing multiseriate eggs; each egg about 180 μm in diameter. Rostrum absent. Antennule (Fig. 117E) about 85 μm long, unsegmented, with traces of 3 weak articulations along posterior side; armed with 15 setae (3 distal setae larger than others). Antenna (Fig. 117F) broad, 3-segment- ed; first segment unarmed; second segment wider than long, with short outer and longer inner margins, and with 1 seta at inner distal corner; third segment 1.5 times longer than wide, with 1 setiform process, 2 setae (including very small one) and 1 spinulose lobe. Labrum (Fig. 117G) much wider than long, unornamented, with concave posterior margin. Mandible (Fig. 117H) consisting of segment plus strong, straight distal spine bearing 1 row of spinules distally. Maxillule absent. Maxilla (Fig. 117I) unsegmented, but divisible into broad proximal and narrower distal parts; armed with 2 setae (1 apical and 1 subapical). Maxilliped (Fig. 117J) broad, 2-segmented; proximal segment very broad, unarmed; distal segment with several patches of minute spinules on posterior surface (Fig. 117K), bluntly protruded mediodistal corner, and 1 acutely pointed spine on anteromedial surface; terminal claw with bicuspid tip. Leg 1 (Fig. 117L) consisting of protopod, exopod, and endopod. Protopod bearing 1 small seta on outer margin, 1 short tubercle on posterior margin between rami, and 4 tubercles at inner distal corner. Exopod with bifurcate anterior lobe and rounded posterior lobe lacking distal seta. Endopod wider than long, with 1 large tubercle on anterior surface. Ventral lateral surface of body lateral to leg 1 bearing 1 tubercle. Ventral surface of body between left and right legs 1 bearing 10 rounded interpodal tubercles (replacing intercoxal plate). Leg 2 (Fig. 118A) similar to leg 1, except bearing 2 distal tubercles on endopod (large outer and small inner). Leg 3 same as leg 2. Leg 4 (Fig. 118B) similar to leg 2, except bearing 9 interpodal tubercles and 2 small tubercles at inner distal corner of endopod. Leg 5 (Fig. 118C) as tapering, rudimentary lobe, wider than long, tipped with 2 small setae. Male. Unknown. Remarks. eaplçstçmella multẚnçdçsa sp. nov. is a very distinctive species, readily distinguishable from all of its congeners by the possession of the characteristic, multiple interpodal tubercles between left and right members of legs 1-4 of the female. These can be observed without dissection and it is likely these tubercles are derived from the transformed intercoxal plate. Tubercles are also present at the inner distal corner of protopods (invariably 4) and on the endopods (1 on leg 1 and 2 on legs 2-4). As additional diagnostic features of the new species, the ventral body surface lateral to each of legs 1-4 bears a single tubercle (a feature shared with e. unẚserẚata sp. nov.), the mandible bears a strong spine, and the body is unsegmented, lacking any trace of segmentation.
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How this classification was reachedexpand
Full frame machine prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. The Gemma side is a direct model label for every work in the frame, read from the title-only record. The Codex side is a classifier learned from the 10,348 direct Codex labels and calibrated to design-weighted sample rates; fields without enough sample support carry no Codex call. Candidate is the union of the two sides; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels.
Distilled classifier scores by category (both heads)
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.000 | 0.001 |
| Meta-epidemiology (narrow) | 0.001 | 0.000 |
| Meta-epidemiology (broad) | 0.001 | 0.000 |
| Bibliometrics | 0.002 | 0.002 |
| Science and technology studies | 0.002 | 0.001 |
| Scholarly communication | 0.001 | 0.002 |
| Open science | 0.001 | 0.001 |
| Research integrity | 0.001 | 0.001 |
| Insufficient payload (model declined to judge) | 0.014 | 0.010 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from itClassification
machine, unvalidatedMachine predicted; a candidate call from one source (direct Gemma or distilled Codex), not a consensus.
How this classification was reached, model by model and score by score, is at the end of the page under "How this classification was reached".