Scipopus (Scipopus) nigripennis GMPAC

Scipopus (Scipopus) nigripennis (Hendel, 1922) Figs 26–27 Tanypoda nigripennis Hendel, 1922: 321. Scipopus albimanus Enderlein, 1922: 212. Scipopus limbativertex Enderlein, 1922: 211. Syn. nov. Scipopus limbativertex – Frey 1927: 74 (listed). — Cresson 1930: 328 (diagnosis). — Hennig 1934: 328 (key). — Aczél 1949: 342 (catalog); 1951: 539 (key). — Steyskal 1968: 48.16 (catalog). — Schumann 1988: 99 (catalog). — Ferro & de Carvalho 2014: 60 (listed). — Marshall et al. 2016: 544 (catalog). Scipopus albimanus – Hennig 1934: 328 (synonymized with S. nigripennis). — Schumann 1988: 85 (catalog). Scipopus nigripennis – Hennig 1934: 328 (key). — Aczél 1949: 342 (catalog). — Steyskal 1968: 48.16 (catalog). — Marshall et al. 2016: 544 (catalog). Differential diagnosis Scipopus (Scipopus) nigripennis resembles S. (S.) furcifer in having an at least partially white first fore tarsomere and a shiny epicephalon that is clearly delineated from the upper frontal vitta, but it differs by the white microtrichose ventral corner of the katepisternum. Type material examined Holotype (Tanypoda nigripennis, photos provided by Peter Sehnal at NHMW, Oct 2020) ECUADOR • 1 ♀; Balzapamba; R. Haensch leg.; NHMW. Syntypes (Scipopus albimanus, not examined) COLOMBIA • 1 ♂, 1 ♀; “Cordilliere, terra caliente”; Dr. Thieme leg.; MNBG. Syntypes (Scipopus limbativertex, examined and photographed in Berlin, 2002, additional photos provided by Jenny Pohl, MNBG) COLOMBIA • 1 ♂; Cordillieren (terra caliente); Dr Thieme leg.; MNBG. BRAZIL • 1 ♀; Matto Grosso; Rhode leg.; MNBG. COLOMBIA • 2 ♂♂; Cordillieren (terra caliente); Dr Thieme leg.; MNBG • 1 ♀; Moritz leg.; MNBG • 1 ♀; Muzo; Rhode leg.; MNBG. MEXICO • 1 ♀; Soconusco; 30 Jul. 1911; C.A. Purpus leg.; MNBG • 1 ♀; same collection data as for preceding; 22 Jul. 1911; MNBG. Other material examined COSTA RICA • 1 ♂, 2 ♀♀; Heredia, Santo Domingo, INBio Pk.; 19–21 Aug. 2001; S.A. Marshall leg.; DEBU • 1 ♀; same collection data as for preceding; USNM • 1 ♀; Monteverde Cloud Forest; 1500 m a.s.l.; 21–26 Aug. 1993; E.R. Barr leg.; Malaise traps; MNCR • 1 ♀; Turrialba; 15 Jul. 1965; P.J. Spangler leg.; USNM • 1 ♂; San Jose, Farm La Caja; 15 May 1919; H. Schmidt leg.; USNM • 1 ♂; Prov. Puntarenas, 2 km N of Hacienda Barú de Dominical Reserve; 22–25 Feb. 2008, S.A. Marshall leg.; MNCR • 1 ♂; same collection data as for preceding; debu00294649/MYCRO545-19 sequenced for CO1–5′; DEBU • 1 ♂; same collection data as for preceding; debu00294650/MYCRO555-19 sequenced for CO1–5′, CO1–3′, 12S, 28S; DEBU • 1 ♂; Prov. Puntarenas, Golfito, P.N. Corvocado, Sector Los Patos, S of al Mirador; 130 m a.s.l.; 15 Nov. 2000; K. Caballero leg.; manual; L_S_517200_279665#61791; MNCR • 1?; same collection data as for preceding; UGIC261-15/MYCRO162-15 sequenced for CO1– 5′; L_S_517200_279665#61792; MNCR • 1 ♀; Alaljuela, Volcán Tenorio, N slope nr Bijagua Biol. Stn; 700 m a.s.l.; 19 Jun. 2000; M. Buck leg.; rain forest; MNCR • 1 ♀; Puntarenas, Corcovado Natl. Pk., San Pedrillo; 8°37′15″ N, 83°44′6″ W]; 5–50 m a.s.l.; M. Buck leg.; human dung; UGIC129-15/ MYCRO030-15 sequenced for CO1–5′; DEBU • 1 ♀; San José, San Carlos, Riosparaiso Res., Pecari Stn, 16 km NNE of Quepos; 9°33′53″ N, 84°7′32″ W; 400 m a.s.l.; 15 Feb. 2003; S.A. Marshall leg.; MNCR • 4 ♀♀; Puntarenas, Osa Peninsula, 2.5 km D of Rincón; 8°42′1″ N, 83°30′50″ S; ~ 50 m a.s.l.; 10–11 Aug. 2001; S.A. Marshall leg.; MNCR (dissected and photographed, Fig. 26C) • 1 ♂, 1 ♀; Alajuela, Volcán Tenorio, N slope nr Bijagua Biol. Stn; 700 m a.s.l.; 17 Jun. 2000; M. Buck leg.; rain forest; MNCR (dissected and photographed, Fig. 26B, E) • 1 ♀; same collection data as for preceding; 16–20 Jun. 2000; S.A. Marshall leg.; RET over Atta mound; DEBU • 1 ♀; same collection data as for preceding; rain forest; UGIC132-15/MYCRO033-15 sequenced for CO1–5′; DEBU • 2 ♀♀; same collection data as for preceding; trail to laguna; ~ 1000 m a.s.l.; rain forest; DEBU • 2 ♀♀; same collection data as for preceding; 19 Jun. 2000; Buck and Marshall leg.; pan traps in tree fall; MNCR • 1 ♀; same collection data as for preceding; debu00138141/MYCRO514-19 sequenced for CO1–5′; DEBU • 1 ♀; Alajuela, San Ramon Biol. Res.; 900 m a.s.l.; May 2000; P. Hanson leg.; MNCR • 1 ♀; San Juan de Peñas Blancas, San Ramón, Soltis Research Centre; 17 Jul. 2013; S.A. Marshall leg.; debu01088995/ MYCRO547-19 sequenced for CO1–5′, CO1–3′, 12S, 28S; DEBU (photographed, Fig. 26G) • 2 ♀♀; Alajuela, San Gerardo Biol. Stn; 10°52′51″ N, 85°23′20″ W; ~ 590 m a.s.l.; 15 Aug 2010; M.D. Jackson leg.; MNCR (photographed, Fig. 26F) • 1 ♀; San José, San Carlos Riosparaiso Res. Pecari Stn, 16 km NNE of Quepos; 9°33′53″ N, 84°7′32″ W; 400 m a.sl.; 12–15 Apr. 2006; S.A. Marshall leg.; DEBU • 1 ♀; same locality as for preceding, upper trail to Rio Negro; 26 Feb. 2006; S.M. Paiero leg.; 1° forest; MNCR • 1 ♀; same locality as for preceding, along road from lodge; ~ 1000 m a.s.l.; 15 Apr. 2015; S.A. Marshall leg.; debu00260110/MYCRO546-19 sequenced for CO1–5′; DEBU • 1 ♀; San Jose, Heredia Biodiversity Garden; 2 Jun. 1998; S.A. Marshall leg.; DEBU • 1 ♀; Guanacaste, Santa Rosa Natl. Pk., hilltop nr La Casona; 10°50′3″ N, 85°36′41″ W; 320 m a.s.l.; 16 Aug. 2016; M.D. Jackson leg.; DEBU • 2 ♂♂, 2 ♀♀, 2?; Heredia, Santo Domingo, INBio Pk.; 19–21 Aug. 2001; S.A. Marshall leg.; DEBU • 3 ♂♂, 1 ♀; same locality as for preceding; 3 Oct. 1999; Marshall and Buck leg.; DEBU • 1 ♀; same locality as for preceding; Jun. 2000; M. Buck leg.; DEBU • 1 ♀; Heredia, etac. Biol. La Selva; 2 Apr. 1993; ALAS exped.; Malaise trap; M/11/058; MNCR • 1 ♀; same locality as for preceding; 2 Mar. 1993; ALAS exped.; MNCR • 1 ♀; Cartago, El Copal. Biol. Res., along Gárvula trail; 9°47′ N, 83°45′ W; ~ 1100 m a.s.l.; 4–7 Aug. 2010; G.F.G. Miranda leg.; DEBU • 1 ♀; Cartago Prov., Turriabla; 13 Jul. 1965; G. and K. Eickwort leg.; SEMC • 1 ♀, 1?; same locality as for preceding; “4” Jun. 1951; O.L. Cartworth leg.; USNM • 1?; same collection data as for preceding, 6 Jun. 1951; USNM • 1 ♀; same collection data as for preceding; 22 May 1951; USNM. HONDURAS • 1 ♂; Gracias a Dios, Rio Patuca, Krausirpe; 22–24 May 1994; B.D. Gill leg.; debu00138356/MYCRO563-19 unsuccessfully sequenced for CO1–5′; DEBU • 1 ♀; same collection data as for preceding; debu00138349/MYCRO644-20 unsuccessfully sequenced for CO1–5′; DEBU. PANAMA • 1 ♂; Potrerillos; 27 Apr. 1954; D.V. Brown leg.; USNM • 1 ♂; Barro Colo. C.Z.; 22 Apr. 1929; S.W. Frost leg.; USNM • 1 ♂; same collection data as for preceding; 12 Feb. 1929; USNM • 1 ♂; same collection data as for preceding; 1 Apr. 1929; USNM • 1 ♀; same collection data as for preceding; Apr. 16 1926; USNM • 1 ♀; Canal Zone Barro Colorado Island; 25 Mar. 1956; C.W. and M.E. Rettenmeyer leg.; SEMC • 1 ♀; Summit, C.Z.; Nov. 1946; N.L.H. Krauss leg.; USNM • 1 ♀; same collection data as for preceding; Oct. 1946; USNM • 1 ♂; Barro Colorado Isl. Canal Zone; 11 May 1978; Silberglied and Aiello leg.; DEBU • 1 ♀; Barro Colorado Isl.; 1–9 May 1964; W.D. and S.S. Duckworth leg.; USNM. Description LENGTH. 11–15 mm. HEAD. Palpus dark brown, pale brown microtrichose and setulose on entire surface, microtrichosity denser on ventral edge and apex, narrow (length 5.0× height). Clypeus light brown to black, width 1.2–2.0 × height, bare medially, white microtrichose posterolaterally. Frontal vitta and orbital plate dull, orange, microtrichose. Epicephalon light to dark brown, shiny, mostly bare, white microtrichose near upper fronto-orbital, narrow (width <⅔ of upper frontal vitta width at upper fronto-orbitals), clearly delineated. Paracephalon light to dark brown, slightly convex on outer portions. All head chaeototaxy well-developed. THORAX. Scutum black-brown, light brown microtrichose, with narrow, anteromedian pale brown vitta flanked by black-brown vittae overlayed with a blue sheen; dull silvery-blue patch anterolaterally to transverse suture. Cervical sclerite slightly convex. Postpronotal lobe black-brown, silvery microtrichose anterodorsally, some black setulae on outer lateral margin. Notopleuron black-brown, pale microtrichose line or spot posterior to anterior notopleural seta extending onto ventral edge. Pleuron with the following microtrichosity: entirely white on proepisternum, almost entirely white on proepimeron, with brown on posterodorsal corner, white on anteroventral ½ and posterodorsal corner of anepisternum, sometimes white on dorsal margin (Fig. 27); ventral corner and remainder brown, almost entirely white on katatergite, at some angles small median brown spot visible. Microtrichosity on katepisternum ranges from entirely white with a small median brown spot (Fig. 27A), to large white patches anteriorly and posteriorly, with remainder brown (Figs 26G, 27B). Legs black-brown with blue sheen; first fore tarsomere entirely white dorsally, hind tarsomere 1 almost entirely white dorsally. Wing uniformly black-brown infuscate. ABDOMEN (J+ ♀). T1 with fine, long white setae. Pleuron light grey, microtrichose, P1 and dorsal half of P2–6 dark brown. FEMALE. T1+2 2.0 × length of T3. Oviscape 2.0–3.0 × length of T6, dark brown, sparsely brown setulose, white microtrichose on anterior ⅓. Combined spermathecal ducts long; common duct ~⅔ of total duct length. Paired spermathecal duct long (8–10 × length of paired spermathecae), parallel-sided, slightly swollen apically. Paired spermathecal stems slightly longer than spermathecae, bare or with sparse pointed tubercles. Paired spermathecae approximately spherical. Single spermathecal duct arising from basal ⅛ of paired duct, narrow, <½ diameter and length of paired duct, swollen apically. Single spermatheca apically round and sclerotized, inornate. MALE. T1+2 ~2.1× length of T3. Genital fork ~2.5× length of T6, arms converging, nearly touching, inner basal process straight, ~½ length of arm. Epandrium elongate, (length 2.0× height), taller posteriorly, short setose on posteroventral edge. Basiphallus small, frame-like. Basal dist

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