Pseudopterogramma metatarsalis Kuwahara & Marshall & Luk 2025, comb. nov.
Bibliographic record
Abstract
Pseudopterogramma metatarsalis (Papp, 2008) comb. nov. Figs 65, 68I, 69 Archipterogrammoides metatarsalis Papp, 2008: 87. Archipterogrammoides metatarsalis – Marshall et al. 2011: 234. Type material Holotype THAILAND – Trang • ♂; Khao Pu-Khao Ya National Park; 21 Nov. 2004; L. Papp and M. Földvári leg.; along a forest brook below the (Pak Yam) waterfall; HNHM. Paratypes THAILAND – Trang • 1 ♂; same collection data as for holotype; HNHM • 1 ♀; Khao Chong Botanical Garden; 14 Nov. 2004; L. Papp and M. Földvári leg.; along the stream below waterfall; HNHM • 1 ♂; same collection data as for preceding; 15 Nov. 2004; along a forest path; HNHM • 3 ♂♂, 2 ♀♀; same collection data as for preceding; 18 Nov. 2004; rainforest; HNHM • 7 ♂♂, 4 ♀♀; same collection data as for preceding; 22 Nov. 2004; HNHM • 1 ♀; Thung Khai Botanical Garden; 19 Nov. 2004; L. Papp and M. Földvári leg.; primary lowland rainforest; HNHM. Material examined Holotype photo provided by Petra Szöllősi-Tóth, HNHM. SINGAPORE • 2 ♀♀; Bird Park Buffer (MIS_L08_MT); 2 May 2019; Mandai Insect Survey Project leg.; LKCNHM • 1 ♂, 2 ♀♀; Bird Park Buffer (MIS_L10_MT); 9 May 2019; Mandai Insect Survey Project leg.; DEBU • 2 ♂♂, 3 ♀♀; same collection data as for preceding; LKCNHM • 1 ♂, 1 ♀; Quarantine Area at Zoo (MIS_L05_MT); 30 May 2019; Mandai Insect Survey Project leg.; LKCNHM. Redescription BODY (Fig. 65). Length 1.0– 1.5 mm. Head yellow, darkened medially. Interfrontal bristles in three pairs. Antennal scape with an enlarged anterior seta. Thorax yellow-brown, scutum caramel-brown. Anterior dorsocentral bristles separated by 7–8 rows of acrostichal setulae. Inner postpronotal seta minute. Femora yellow-brown, tibiae brown with broad yellow medial band. Ventral surface of male mid tibia with a long apicoventral bristle. Mid basitarsus with a slightly enlarged ventral setula. Hind tibia with a stout apicoventral spur. Wing (Fig. 68I) infuscate with several pale vertical blotches. R 2+3 almost straight. MALE ABDOMEN. S5 strongly asymmetrical, the left side much longer than the right, with a broad posteromedial emargination. S6+7 well-developed; apical third of S6 large and triangular, with a distinct constriction between it and the basal two-thirds, left third of S6 with a medially-projecting triangular lobe. Cercus low and semicircular with two long and several short setae; subepandrial sclerite broad, well-developed, U-shaped with a pair of thin connections to the surstylus. Anterior apodeme of hypandrium short and broad, lateral arms broad. Surstylus short, ovoid, relatively sparsely setose except subbasally and ventrally, with a thickened, tooth-like ventromedial seta. Postgonite elongate, thin, slightly sinuate and apically truncate. Phallapodeme elongate, narrow with a slightly enlarged apex; basiphallus large, thick, and tubular, slightly curved with an anterobasal lip and a subapical notch where the distiphallus articulates. Distiphallus large, “conspicuously empty”, and composed of several sclerites: a long, slightly tapered ventral sclerite with a downturned, bifurcate apex, a pair of S-shaped, subventral sclerites articulating with the apicolateral corners of the ventral sclerite, a broad, saddle-shaped posterodorsal sclerite, and an elongate, apically bifurcate, sinuate sclerite just ventral to the dorsal sclerite; the apex of the distiphallus is membranous between these sclerites. FEMALE ABDOMEN. T2–4 reduced, leaving a narrow membranous area between T2 and T3, and a larger membranous area between T3 and T4. Epiproct and hypoproct small and weakly sclerotized. Cercus short, broad, and often angled upwards, with long apical, subapical, and lateral setae. Spermathecae stout, bowl-shaped, stem with a globular base, short in the single spermatheca and long in the paired spermathecae. Distribution Indomalayan: Singapore (new record), Thailand (Papp 2008). Remarks Several characters of the male genitalia (long, sinuate postgonite; large basiphallus roughly half the length of the distiphallus; distiphallus composed of many well-developed sclerites) seem to be synapomorphic with Ps. aestivale, Ps. brevivenosum, and Ps. conicum, suggesting that the treatment of metatarsalis as a separate genus Archipterogrammoides rendered Pseudopterogramma paraphyletic.
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How this classification was reachedexpand
Full frame machine prediction
Teacher imitationNot calibrated prevalence, not ground truth. Human validation pending. The Gemma side is a direct model label for every work in the frame, read from the title-only record. The Codex side is a classifier learned from the 10,348 direct Codex labels and calibrated to design-weighted sample rates; fields without enough sample support carry no Codex call. Candidate is the union of the two sides; consensus is their intersection. These outputs are machine_predicted_unvalidated and are not human labels.
Distilled classifier scores by category (both heads)
| Category | Codex | Gemma |
|---|---|---|
| Metaresearch | 0.000 | 0.001 |
| Meta-epidemiology (narrow) | 0.001 | 0.000 |
| Meta-epidemiology (broad) | 0.000 | 0.000 |
| Bibliometrics | 0.001 | 0.001 |
| Science and technology studies | 0.002 | 0.001 |
| Scholarly communication | 0.000 | 0.002 |
| Open science | 0.001 | 0.001 |
| Research integrity | 0.001 | 0.001 |
| Insufficient payload (model declined to judge) | 0.015 | 0.011 |
Machine scores (provisional)
The two teacher heads of the student model, read on this work. A score orders the frame for review; it never asserts a category, and the validation status ships verbatim with every row.
Baseline scores from an immature model (maturity gate not passed, 7 training rounds). Scores rank; they never assert a category.
score_only:v0-immature-baseline · verbatim from the scoring run: score_only means the number may rank works, and no category label ships from itClassification
machine, unvalidatedMachine predicted; a candidate call from one source (direct Gemma or distilled Codex), not a consensus.
How this classification was reached, model by model and score by score, is at the end of the page under "How this classification was reached".