Atheta graminicola

Atheta (s. str.) graminicola (Gravenhorst, 1806) (Fig. 155 in Strand & Vik 1964) Aleochara graminicola Gravenhorst, 1806: 176. Homalota granulata Mannerheim, 1846: 508 (misspelled as Stonalota granulata). Liogluta (Megista) graminicola: Mulsant & Rey, 1874a: 624 (as valid species). Atheta (Megista) graminicola: Bernhauer, 1907: 394 (as valid species). Atheta graminicola: Fenyes, 1907: 61 (as valid species). Atheta granulata: Fenyes, 1907: 61 (as synonym of At. graminicola). Atheta (Homalotusa) lanei Casey, 1910a: 10, syn. nov. Atheta (Megista) nomadica Casey, 1910a: 15, syn. nov. Atheta (Megista) graminicola: Bernhauer & Scheerpeltz, 1926: 659 (as valid species). Atheta (Megista) granulata: Bernhauer & Scheerpeltz, 1926: 659 (as synonym of At. graminicola). Atheta (Megista) nomadica: Bernhauer & Scheerpeltz, 1926: 659 (as valid species). Atheta (Homalotusa) lanei: Bernhauer & Scheerpeltz, 1926: 655 (as valid species). Atheta (s. str.) graminicola: Strand & Vik, 1964: 155 (as valid species). Atheta (s. str.) graminicola: Benick & Lohse, 1974: 195 (as valid species). Atheta graminicola: Lohse & Smetana, 1985: 286 (as valid species). Atheta granulata: Lohse & Smetana, 1985: 284 (as valid species). Atheta graminicola: Lohse et al., 1990: 190 (as valid species). Atheta granulata: Lohse et al., 1990: 189 (as valid species). Atheta nomadica: Lohse et al., 1990: 189 (as synonym of At. granulata). (Other references are omitted) Type material. Lectotype of Aleochara graminicola (here designated): ♂, “5349”, “ graminicola Gr. ”, “Zool. Mus. Berlin ” (ZMHB). Paralectotypes: 2♂♂, 2♀♀, without labels; ♀, “Angl. Rudd”; ♀, “Nordeur. Schupp.” (ZMHB). Lectotype of Homalota granulata (designated by Lohse and Smetana (1985)): UNITED STATES: Alaska: ♀, Unalaska Island (Cygnaeus) (MZHF). Lectotype of Atheta lanei (here designated): ♀, “ Lane Co., Or.[egon]”, “ lanei Csy. ”, “TYPE USNM 39462” (red label), “CASEY bequest 1925” (NMNH). Lectotype of Atheta nomadica (here designated): ♀, “ Nome, Alaska ”, “ nomadica Csy. ”, “TYPE USNM 39478” (red label), “CASEY bequest 1925” (NMNH). Additional material. CANADA: Alberta: ♂, ♀, Banff (Fenyes); 3♀♀, 2 mi. S Ponaka, Battle Riv. marsh, 6.viii.1976 (P.M.Hammond); ♂, 2♀♀, Waterton N. P., Cameron Lake, 540’, 2–4.viii.1976 (P.M.Hammond) (BMNH); British Columbia: ♂, Alta Lake, 1.viii.1975 (J.M. & B.A.Campbell) (CNCI); Labrador: ♀, L’Anse au Loup, 9.viii.1972 (J.M.Campbell); 2♂♂, Hebron, 9.vii.1954 (J.F.McAlpine) (CNCI); Manitoba: ♀, Churchill, 29.vi.1937 (W.J.Brown); ♀, ditto but 20.vii.1937; ♂, ditto but 18.viii.1937; ♂, 2♀♀, ditto but 16.vi.1937; 2♂♂, 6♀♀, ditto but 4.vi.1952 (J.G.Chillcott); 4♂♂, 4♀♀, ditto but 13.vi.1952; ♂, 3♀♀, ditto but 17.vi.1952; ♂, ♀, ditto but 2.viii.1952; 3♂♂, ♀, ditto but 5.viii.1952; ♂, ditto but 15.viii.1952; ♀, ditto but 27.viii.1948 (W.R.Richards) (CNCI); Newfoundland: ♂, Long Range Mts., 50º34’N 56º56’W, 1400’, 24.vii.1969 (M.J.D.Brendell) (BMNH); Northwest Territories: 2♀♀, Ft. Simpson, Harris River, 15.vi.1972 (A.Smetana); ♂, ditto but 10.vi.1972; 5♂♂, 19♀♀, Trail River, 32 mi. NW Ft. Simpson, 16.vi.1972 (A.Smetana); 3♂♂, 4♀♀, Hwy. 3, 5 mi. SE Ft. Simpson, 17.vi.1972 (A.Smetana); 3♂♂, 2♀♀, ditto but 21.vi.1972; ♀, Hwy. 3, 8 mi. SE Ft. Simpson, 18.vi.1972 (A.Smetana); ♂, Spence River, 38 mi. SE Ft. Simpson, 19.vi.1972 (A.Smetana); 3♂♂, 13♀♀, Reindeer Sta., Caribou Hills, 30.vi.1972 (A.Smetana); ♀, Yellowknife, 20.v.1953 (J.G.Chillcott); ♂, ditto but 29.v.1953; 2♀♀, Aklavik, 9.vi.1956 (R.E.Leech); 2♀♀, ditto but 12.vi.1956; ♂, ditto but 14.vi.1956; ♂, ditto but 3.vii.1956; ♀, ditto but 6.vii.1956; ♂, ditto but 13.vi.1956 (E.F.Cashman); ♀, ditto but 16.vi.1953 (C.D.Bird); 10♀♀, Inuvik, Shell Lk., 27.vi.1972 (A.Smetana); ♀, ditto but 1.vii.1972 (A.Smetana); 6♀♀, ditto but 25.vi.1972; 2♂♂, ♀, Reindeer Depot, Mackenzie Delta, 8.viii.1948 (W.J.Brown); ♀, Kidluit Bay, N. Richards Is., 26.vii.1948 (T.J.R.Vockeroth); ♀, Muscox L., 64º45’N 108º10’W, 2.viii.1953 (J.G.Chillcott); 3♂♂, 1 specimen, ditto but 21.vii.1953 (J.G.Chillcott); ♂, Salmita Mines, 64º05’N 111º15’W, 20.vi.1953 (J.G.Chillcott); 2♀♀, Unnamed Lake, 18 mi. NW Inuvik via East Channel, 26.vi.1972 (A.Smetana) (CNCI); Ontario: ♂, 2♀♀, Butterfly Lk., 13 mi. S Sioux Lookout, 17.vi.1973 (Campbell & Parry) (CNCI); Québec: 7♂♂, 15♀♀, Blanc Sablon, 9.viii.1972 (J.M.Campbell); ♂, 2♀♀, Koroc Riv., in leaf litter beside stream, 3.viii.1986 (A.V.A. & S. Morgan); ♂, Whale Riv., 8.vii.1949 (J.R.Vockeroth) (CNCI); Yukon Territory: ♀, Caribou Bar Creek, 20.vi.1972; 2♂♂, 5♀♀, McQuesten Lk., 15 mi. SW Keno, 19.vii.1968 (Campbell & Smetana); 2♂♂, 3♀♀, Mi. 1034, Alaska Hwy., near Kloo Lake, 5.vii.1968 (Campbell & Smetana); 4♀♀, Mi. 1120, Alaska Hwy., 6.vii.1968 (Campbell & Smetana); ♀, Mi. 1192, Alaska Hwy., near Snag Junction, 6.vii.1968 (Campbell & Smetana); ♀, Mi. 1120, Alaska Hwy., 6.vii.1968 (Campbell & Smetana); ♂, ♀, Mi. 1705, Alaska Hwy., Slim’s River, 7.vii.1981 (Bright); ♂, 10 mi. E Dawson City, 11.vii.1968 (Campbell & Smetana); 3♂♂, 2♀♀, Dempster Hwy., Mi. 236, Eagle Riv., 1400’, 12.vii.1978 (Campbell & Smetana); 2♂♂, Dempster Hwy., Mi. 222, 1900’, 21.vii.1978 (Smetana & Campbell); 3♂♂, 6♀♀, ditto but Mi. 147, 22.vii.1978 (Campbell & Smetana); 2♀♀, ditto but Mi. 139.5, 20.vii.1978 (Campbell & Smetana); ♀, ditto but Mi. 73, 3300’, 12.vii.1968 (Campbell & Smetana); 2♂♂, 10♀♀, ditto but Mi. 71; ♂, 7♀♀, ditto but Mi. 65, 3300’, 13.vii.1968; 4♂♂, ♀, ditto but Mi. 60, 3500’, 19.vii.1978; 4♂♂, 4♀♀, ditto but 23.vii.1978; ♂, 2♀♀, ditto but Mi. 55, 3300’, 13.vii.1968; 5♂♂, 2♀♀, ditto but Mi. 53, North Fork Pass, 4200’, 24.vii.1978; 3♂♂, ♀, ditto but Mi. 48.5, North Fork Pass, 4100’, 24.vii.1978; ♂, 3♀♀, ditto but 19.vii.1978; 5♂♂, 2♀♀, ditto but Mi. 45, 3500’, 14.vii.1968; ♀, ditto but 3400’, 18–23.vii.1978 (J.M.Campbell); 11♂♂, 13♀♀, ditto but Mi. 42, N Klondike Riv., 3300’, 18.vii.1978 (Campbell & Smetana); ♂, ditto but Mi. 32, 2700’, 15.vii.1968; ♂, ditto but Mi. 28, 15.vii.1968; ♀, Ogilvie Mts., North Fork Pass, 4100’, 18.vi.1962 (P.J.Skitsko); ♂, ♀, Rampart House, 28.v.1951 (J.E.H.Martin); ♂, ditto but 1.vi.1951; 4♀♀, British Mts., 340 m, 4 km NE Wolf Cr., 69º08’N 140º14’W, 23.vi.1984 (Campbell); ♂, ♀, British Mts., 320 m, 3 km NW Firth Riv., 69º13’N 140º05’W, sifting Salix litter, 19.vi.1984 (Campbell); ♂, ♀, British Mts., 180 m, Fish Creek, 69º27’N 140º19’W, sifting under tall Salix shrubs, 3.vii.1984 (J.M.Campbell); 2♂♂, 3♀♀, Hansen Lk., 9 mi. SW Keno, 18.vii.1968 (Campbell & Smetana); ♂, La Force Lake, 62º41’N 132º20’W, 3300’, 28.vi.1960 (J.E.H.Martin); ♀, Swim Lakes, 62º13’N 133º00’W, 3200’, 15.vi.1960 (J.E.H.Martin) (CNCI); ♀, Whitehorse, 21.vi.1924 (MCZ); UNITED STATES: Alaska: 16♂♂, 17♀♀, Mi. 123, Taylor Hwy., 9.vii.1968 (Campbell & Smetana); ♀, Mi. 23, Taylor Hwy., 8.vii.1968 (Campbell & Smetana); ♀, Denali State Park, Mi. 165.5, Hwy. 3, 23.vi.1978 (Smetana & Becker); 2♂♂, 3♀♀, Mi. 110, Denali Hwy., Seattle Cr., 15.vii.1978 (Campbell & Smetana); 4♂♂, 5♀♀, Mi. 102, Denali Hwy., 12.vii.1978 (Smetana & Campbell); 2♀♀, Mi. 94, Denali Hwy., Canyon Creek, 13.vii.1978 (J.M.Campbell & A.Smetana); ♂, 12 mi. N Mi. 78, Denali Hwy., Windy Cr., 13.vii.1978 (Smetana & Campbell); 6♂♂, 8♀♀, Mi. 1249, Alaska Hwy., Deadman Lk., 6–7.vii.1968 (Campbell & Smetana); 2♂♂, 6♀♀, Mi. 1252, Alaska Hwy., 7.vii.1968 (Campbell & Smetana); 2♂♂, ♀, Kenai Pen., Kenai, 24.vi.1951 (W.J.Brown); ♂, ♀, Kenai Pen., 3 mi. SE Kenai, 7.vi.1978 (Smetana & Becker); ♀, Kenai Pen., Hope, 24.v.1951 (W.J.Brown); 5♂♂, 7♀♀, Kenai Pen., Anchor River at Hwy. 1, 450’, 2.vi.1978 (Smetana & Becker); ♂, 20♀♀, ditto but 4.vi.1978; ♀, ditto but 5.vi.1978; ♂, 4♀♀, Kenai Pen., Anchor River Campground, 12 mi. N Homer, 450’, 5.vi.1978 (Smetana & Becker); ♀, ditto but 3.vi.1978; 4♂♂, 5♀♀, Kenai Pen., 1 mi. W Anchor Pt., 1.vi.1978 (Smetana & Becker); 7♂♂, 3♀♀, Kenai Pen., Crooked Cr. at Hwy. 1, S Kasilof, 9.vi.1978 (Smetana & Becker); 5♂♂, 11♀♀, Kenai Mts., 15 mi. N Seward, 400’, 29.v.1978 (Smetana & Becker); 6♂♂, 9♀♀, Kenai Mts., 16 mi. N Seward, 500’–600’, 26.v.1978 (Smetana & Becker); 4♂♂, 6♀♀, Kenai Mts., 22 mi. N Seward, 600’–800’, 27.v.1978 (Smetana & Becker); 2♂♂, 5♀♀, Seward, 20.v.1951 (W.J.Brown); 2♂♂, Kenai Pen., 15 mi. E Sterling, 14.vi.1978 (Smetana & Campbell); 2♂♂, ♀, Kenai Pen., 2 mi. NE Soldotna, 10.vi.1978 (Smetana & Becker); 4♂♂, Kenai Pen., Clam Gulch, 6.vi.1978 (Smetana & Becker); 3♂♂, 5♀♀, Kenai Mts., Cooper Lk., 1000’, 15.vi.1978 (Smetana & Becker); ♀, Kenai Mts., 8 mi. W Cooper Landing, 500’, 14.vi.1978 (Smetana & Becker); ♀, Kenai Mts., Quarts Cr. At Hwy. 1, 500’, 15.vi.1978 (Smetana & Becker); ♂, 3♀♀, Kenai Mts., Tern Lk. Campground, 700’, 18.vi.1978 (Smetana & Becker); ♀, 8 mi. NW Haines, 3.vii.1968 (Campbell & Smetana) (CNCI); 3 specimens, Skagway, 18– 20.vi.1924 (H.C.Fall) (CUIC); 4♂♂, ♀, ditto but 18.vi.1924 (MCZ); 3♂♂, 8♀♀, Hess Cr., Mi. 24, Wales Hwy., 1.vii.1978 (J.M.Campbell & A.Smetana); 3♂♂, ♀, ditto but 600’, 65º40’N 149º10’W, 10.vii.1978 (J.M.Campbell & A.Smetana); 9♂♂, 5♀♀, Unalakleet, 4.vii.1961 (R.Madge); 2♀♀, ditto but 23.vi.1961; ♀, ditto but 17.vi.1961; ♀, ditto but 11.viii.1961; ♀, King Salmon, Naknek River, 6.vii.1952 (W.R.Mason); 1 specimen, Nome, 10.vii.1951 (D.P.Whillians); ♂, Mi. 220, George Parks Hwy., 14.vii.1978 (A.Smetana) (CNCI); 2♀♀, Fairbanks, 7.vii.1924 (MCZ); 2♀♀, Prudhoe Bay Rd., Bonanze Cr., 66º40’N 150º40’W, 900’, 2.vii.1978 (A.Smetana & J.M.Campbell); ♀, Prudhoe Bay Rd., 4 mi. SW Chandalar Cmp., 68º04’N 149º40’W, 4000’, 7.vii.1978 (Campbell & Smetana); 2♂♂, 4♀♀, Prudhoe Bay Rd., Coldfoot nr. Clara Cr., 67º16’N 150º10’W, 1100’, 3.vii.1978 (A.Smetana & J.M.Campbell); 3♀♀, Prudhoe Bay Rd., 8 mi. N S. Fork Koyukuk Riv., 67º13’N 150º07’W, 1000’, 8.vii.1978 (A.Smetana & J.M.Campbell); 4♂♂, 6♀♀, Prudhoe Bay Rd., 0.5 mi. N Yukon Riv., 65º52’N 149º45’W, 400’, 9.vii.1978 (A.Smetana & J.M.Campbell) (CNCI); 2 specimens, Glenn Hwy. (Tok cut­off), NE of Gakona, 62º19.89–.88'N 145º12.60–.27'W, 550 m, a bog, lake shore, Carex, C

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