Quedius fuliginosus

Quedius fuliginosus (Gravenhorst, 1802) (Figs 1, 2A,E, 3, 6C, 7C, 11D, 12, 18) Staphylinus fuliginosus Gravenhorst, 1802: 34 [Type locality:Brunsviga]. Note: See lectotype designation by Gඎඌൺඋඈඏ (1993) Quedius granulipennis Motschulsky, 1858: 656 [Type locality: Autriche]. Note: See lectotype designation by Gඎඌൺඋඈඏ (1993) Quedius impunctifrons Delahon, 1915: 395. [Type locality: Mark Brandenburg] Quedius jelineki Krása, 1904: 81 [Type locality: Vrané n. Vlt, Bohemia] Quedius latus Hochhuth, 1851: 30 [Type locality: Kaukasien, Alagez- Gebirge], syn. nov. Quedius molochinicolor Roubal, 1931: 1 [Type locality:RCS.:Harmanec] Quedius picicornis Stephens, 1832: 215 [Type locality: London] Quedius subfuliginosus Britten, 1944: 290 [Type locality: England: Cotterill Clough, Cheshire] Quedius viduus Sawada, 1965: 17 [Type locality: Japan, Pref. Nagano, Mt. Jônen], syn. nov. References. MൺඋඌHൺආ (1802): 504 (characters); Lൺඍඋൾංඅඅൾ (1804): 319 (characters); GඒඅඅൾඇHൺඅ (1810):301 (as tristis; characters);MൺඇඇൾඋHൾංආ (1830): 25 (distribution); (1831): 439 (distribution); Nඈඋൽආൺඇඇ (1837): 78 (distribution); EඋංർHඌඈඇ (1839a): 490 (distribution); (1840): 537 (characters); Hൾൾඋ (1839): 276 (characters); Dඎൿඈඎඋ (1843): 33 (characters); Kඈඅൾඇൺඍං (1846): 22 (distribution); GඋൺඏൾඇHඈඋඌඍ (1847): 232 (characters); HඈർHHඎඍH (1849): 151 (characters); (1862): 46 (distribution); RൾൽඍൾඇൻൺർHൾඋ (1849): 710, (1857): 203, (1874): 200 (characters); Kංൾඌൾඇඐൾඍඍൾඋ (1851): 419 (distribution); Hൺඋൽඒ (1851): 33 (biology); Kඳඌඍൾඋ (1853): 62 (characters); Fൺංඋආൺංඋൾ & Lൺൻඈඎඅൻජඇൾ (1856): 539 (characters); Kඋൺൺඍඓ (1857): 503 (characters); (1858): 58 (distribution); Pൾඒඋඈඇ (1858): 428 (distribution); Fൺඎඏൾඅ (1865):293 (biology); (1874): 289 (characters); Sൾංൽඅංඍඓ (1875): 267 (characters); Mඎඅඌൺඇඍ & Rൾඒ (1876): 686 (characters); SൺHඅൻൾඋG (1876): 24 (distribution); Fඈඐඅൾඋ (1888): 237 (characters); GൺඇGඅൻൺඎൾඋ (1895): 403 (characters); Pඈඉඉංඎඌ (1905): 9 (distribution); Pඈඋඍൺ (1907): 128 (characters); Rൾංඍඍൾඋ (1909): 111 (characters); JඈHൺඇඌൾඇ (1914): 363 (characters); EංർHൾඅൻൺඎආ (1914): 339, (1915): 91 (morphology); Fංඈඋං (1915): 16 (distribution); Mඎඇඌඍൾඋ (1923): 195 (distribution); Gඋංൽൾඅඅං (1924): 79 (characters); Pඈඋඍൾඏංඇ (1929): 341 (characters); Wඳs ඍHඈൿൿ (1938): fig. 21 (morphology); Pൺඎඅංൺඇ (1941): 263 (larval characters); Hංඇඍඈඇ (1945): 70 (characters); FൺGൾඅ (1948): 196 (characters); Jൾൺඇඇൾඅ & JൺඋඋංGൾ (1949): 378 (biology); Hൺඇඌൾඇ (1952): 140 (characters); Sආൾඍൺඇൺ (1958): 361 (characters and biology); (1962): 133 (characters and distribution); (1964): 79 (distribution); (1976): 22 (distribution); (1978c): 85 (distribution); (1993): 50 (distribution); Pൺඅආ (1963): 139 (characters); LඈHඌൾ (1964):211 (characters); Hඈඋංඈඇ (1965): 272 (distribution); Pඈඍඈඍඌĸൺංൺ (1967): 84, 87 (larval characters); Kൺඌඎඅൾ (1968): 53 (biology); (1970): 63 (larval characters); KඈඋGൾ (1968): 52 (characters); Sඓඎඃൾർĸං (1968): 735 (distribution); Fඋൺඇĸ (1969): 266 (biology); (1982): 46 (list of parasites); Bඈඋൽඈඇං (1974): 12, (1976a): 89 (characters); (1976b): 238 (distribution); Oඌൾඅඅൺ & Zൺඇൾඍඍං (1975): 120 (biology); Pඈඉൾ (1977): 31 (distribution); Cඈංൿൿൺංඍ (1978): 190 (distribution); Tඈඉඉ (1979): 19 (development); Bඎඋൺĸඈඐඌĸං et al. (1980): 134 (distribution); TඬඍH (1984): 117 (characters); Nඈඐඈඌൺൽ (1990):143 (biology);Aඅඅൾඇ (1990): 4 (biology); WHංඍൾHൾൺൽ (1991): 6 (biology); Gඎඌൺඋඈඏ (1991):9 (identity); (1993): 72 (synonymic notes, lectotype designation); WൾඅർH (1993): 229 (morphology); HඈൽGൾ & Jඈඇൾඌ (1995): 42 (characters); Cංർൾඋඈඇං & Zൺඇൾඍඍං (1995):32 (distribution); Sඍൺඇංൾർ (1996):117 (pupal characters); (2010): 22 (distribution); Oඐൾඇ (1997): 149 (biology); Sඍൺඇංൾർ (1999): 52 (characters of pupa); Dൾඋඎඇĸඈඏ (2005): 279 (distribution); UHඅංG et al. (2006): 54 (distribution); Tඋඈඇඊඎൾඍ (2006): 102 (distribution); Mൺඃĸൺ & Sආൾඍൺඇൺ (2007): 428 (distribution); Mൺඓඎඋ et al. (2007): 29 (distribution);Sඍൺඇ (2009):242 (distribution);Sൾආൾඇඈඏ et al.(2015): 127 (distribution); Sൾආංඈඇൾඇĸඈඏ et al. (2015): 331 (distribution); Sൺඅඇංඍඌĸൺ & Sඈඅඈൽඈඏඇංĸඈඏ (2018a): 125, (2019): 48 (distribution); PඎർHĸඈඏ et al. (2020): 43 (distribution). Type material examined. Quedius fuliginosus: Lൾർඍඈඍඒඉൾ J (ZMHB), labelled: “coll. Hellwig / Paratypus fuliginosus Grav. ”. Quedius granulipennis: Lൾർඍඈඍඒඉൾ J (ZMMU), labelled: “ Austria / Quedius granulipennis Motsch. Austria ”. Quedius latus: Nൾඈඍඒඉൾ J (ZMHB), designated here, labelled: “AR-MENIA (AR-16-19) 35 km NW Sisian, 39°40’59”N 45°46’50”E, 2070 m, stream valley, litter beneath bushes near stream sifited 3.VII.2016, leg. M.Schülke / Quedius (Quedius) fuliginosus (Gravenhrost) det. M. Schülke 2019 / Museum für Naturkunde Berlin Sammlung M. Schülke / Neotype Quedius latus Hochhuth, 1851 A.K. Hansen & A. Solodovnikov des. 2021”. Quedius molochinicolor: Sඒඇඍඒඉൾ ♀ (SNMC), labelled: “ Slovensko Harmanec Roubal 26.IX.1930 / f. molochinicolor mi [my] type / [red label] / fuliginosus f. molochinicolor cotypus”. Additional material examined. ARMENIA: N Yerevan, NW Hrazdan, 40.6350 44.4602, grassy W slope with scattered Salix, litter and roots of grass sifted, 2500m, 26.VI.2016, leg. M. Schülke (1 ♀ cSch); same locality, 40.6944, 44.4878, stream valley, mixed decidious forest, litter and grass roots sifted, 2110m, 28.VI.2016, leg. M. Schülke (1 ♀ cSch). AUS-TRIA: Arbesbach; [48.49, 14.95], 1.-13.VII.1951, leg. Schubert (1 ♀ NMW); Brunnlust, Moosbrunn, Schwechat, [48.01, 16.45], 12.VI.1997, leg.Schillhammer (1J NMW);Faak, Kärnten,[46.56, 13.91], 5.VIII.1968, leg. Wewalka (1 ♀ NMW); Immerkrems, [46.96, 13.72], subalpine, 1400 - 2000 m, VI.1954, leg. Lindroth (1 ♀ MZLU); Plesch-Kogel n. Rein, [47.13, 15.28], 590m, sifting forest litter, 20.VI.1995, leg. Zerche & Behne (1 J SDEI); Rekawinkel, [48.17, 16.02], leg. Briet (1 J NMW). BULGARIA: Sofia, Tschernia [Cherni Vrah], [42.56, 23.27], 600m, 22.VI.1988,leg.Behne (1J SDEI); Sredna Gora,S Koprivschtiza, 42.8777, 25.0544, moss under Picea, 1000m, 29.VI.1997, leg. Zerche & Behne (1 ♀ SDEI); Stara Plania, 5km S Ribaritza, 42.75, 24.38, 750m, 7.VI.1997, Beech forest, sifting, leg. Zerche & Behne (1 J SDEI); SW-Pirin, Kurort Popina Laka, 16 km NO Sandanski, [41.56, 23.27], ca. 1350m, Pinus and Picea, under snow remains, sifted, 8.IV.2005 leg. Zerche & Behne (1 J SDEI); W Rhodopen, S Velingrad, Tschernowroh, [41.68, 24.05], 1600m 10.VI.1987, leg. Zerche & Behne (1 ♀ SDEI). CZECH REPUBLIC: Podyjí Nat. Park, forest at river Dyje valley ca. 2.5 km NW of Havraníky, 48.8300, 15.9804, h 240m, sifted flood debris, 8.VI.2016, leg.A. Solodovnikov, J. Jenkins Shaw,A. Hansen (1 ♀ NHMD). DENMARK: Amager, Kalvebod Faelled, Store Høj Søen, 55.6159, 12.5606, sifting leaf litter, debris near lake, 17.III.2019, leg.A. K. Hansen (1 J 1 ♀ NHMD); Lisjerg Forest, 8 km N of Aarhus, 56.2335, 10.1688, h 85m, edges of forest lakes, hand collected in leaf litter and moss, 17.IV.2019, leg.A. K.Hansen (4JJ 4 ♀♀ NHMD); Nors Sø, S of Nors Sø, 57.0197, 8.6197, pitfall trap, dry pine forest, 16-18.VI.2014, leg.Solodovnikov, Hansen, Kraemer, & Justesen (1 J NHMD). GEORGIA: N of Sviri, [41.73, 42.99], pitfall traps, 14.IV.-11.VI.2018, leg. D. Formynikh (1 J cSch); Tusheti, n. Omalo, Abanos Pass, [42.27, 45.51], river valley, 1700m, 14.-17.VII.2016, leg. Heinz (1 J NMEG). GERMANY: Bayern, Eschenlohe, [47.58, 11.18], 3.VIII.1937,leg.Ihssen (1J 1♀ SDEI);Berlin-Blankefelde,[52.62,13.39], 8.V.1994, leg. D. Wrase (1 J SDEI); Biesenthal, [52.76, 13.61], 500m, field sandish-clayish ruderal, 1.IV.2002, leg. D. Wrase (1 ♀ SDEI); Brandenburg Kiesfläche 1km S Oderberg, [52.84, 14.04], 13.X.1990, leg. F. Hieke (1 J SDEI); Brandenburg, Oranienburg, Briesetal, [52.71, 13.31], 12.IV.1992, leg.J. Ziegler (1J SDEI); Dars, Zingst, Freesenbruch, [54.43, 12.68], 20.VII.1987, leg. D. Wrase (1♀ SDEI); Erzgebirge,Prinzgehöhle, [50.63, 12.67], 29.IX.1921, leg. Uhmann (1 ♀ SDEI); Erzgebirge, W Teil, SW Kammweg & Morgenröthe, [50.43, 12.51], 4.V.1990 leg. O. Sorge (1 J SDEI); Finkenkrug n. Berlin, [52.56, 13.03], 12.IV.1898, leg. Dahl (1 J SDEI); Kehlheim Dürnbacher Forst, [48.74, 11.75], (1 ♀ SDEI); Liepnitzsee, n.Wandlitz,[52.74, 13.51], 01.X.1989, leg. M. Uhlig (1J SDEI); Marburg i Hess, [50.81, 8.77], 16.IV.1904, leg. Strand (1 J 1 ♀ SDEI); n. Märkische Buchholz, Königs Wusterhausen, [52.11, 13.76], 12.VI.1980, leg. H.Wendt (1 ♀ SDEI); Martinfeld, Krs. Heiligenstadt, [51.28, 10.18], 21.-29.VI.1990, leg. M. Uhlig (1 J SDEI); München-Grünwald, [48.04, 11.53], 3.VII.1931, leg. Ihssen (1 ♀ SDEI); Nauen, [52.61, 12.87], 19.III.1910, leg.M.Ude (1J SDEI);Sächsischen Schweiz, Rathen,[50.95, 14.08], 1.V.1977, leg.K. H.Mohr (1♀ SDEI); Schwäbisch Gmünd,Röben, [48.79, 9.81], 4.V.1953 (1 J SDEI); Seilershof, Gransee, [53.06, 13.18], 22.V.1988, leg. M. Schülke (1 ♀ SDEI); Suekow, [53.07, 11.79], XI.1974, leg. M. Moritz (3 JJ SDEI); Wandlitz, Liepnitzsee, [52.74, 13.51], 6.I.1981, leg. M. Uhlig (1 J 2 ♀♀ SDEI); Wildeschütz n. Deuben Silbersee Schilfgürtel, [51.12, 12.07], 11.X.1987, leg. M. Uhlig (1 ♀ SDEI). ITALY: Alpie Orobie,Bergamo Castione &Passo delia Presolana, 45.9202, 10.1001, 1170m, 18.-21.VI.2006, leg.P. Schnitter (1 J NMEG); Basilicata, Pognola Ris. L. Pignola, [40.58, 15.74], 700 m, 23.XII.1996, leg. F. Angelini (1 cBor); Trentino, Lago di Tenno,N Riva, [45.93, 10.81], 560m, 2.VII.1994, leg. L. Zerche (1 ♀ SDEI). KAZAKHSTAN: Altai, Bukhtarma riv., Chingistai, [49.19, 85.88], 30.VIII.2010, leg. V.A. Kastcheev (1 J ZIN);Akshatau Mt., NW Ayaguz, Semipalat, [48.25, 79.64], forest leaf litter, 17.VII.1962, leg. L. V.Arnoldi (1 J ZIN); Ivanovsky Mt. Ridge, 32 km S Leninogorsk, [50.04, 83.51], 1300 m, 14.VIII.1986, leg. I. I. Kabak (1 J ZIN). RUSSIA: Aඅඍൺං Kඋൺං: Talmenskij Distr., E of vill. Litvinovka 53.6918, 83.7309, 200m, mixed forest, sifted leaf litter, 24.VI.2019, leg. A. Solodovnikov, A. K. Hansen & A. Tokareva (1 NHMD); Chemalskij Distr., SE of vill. Edigan 51.0546, 86.3791, 1030m, larch-fur forest, sifted leaf litter, 27.VI.2019, leg.A. Solodovnikov & A. K. Hansen (1 NHMD); Chemalsk

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